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Carettochelys insculpta RAMSAY, 1886

IUCN Red List - Carettochelys insculpta - Vulnerable, VU

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Higher TaxaCarettochelyidae, Testudines (turtles) 
Common NamesPig-nosed Turtle, Pitted-Shelled Turtle, Warrajan 
SynonymCarettochelys insculptus RAMSAY 1886 (error typographicus)
Carettochelys insculpta — BOULENGER 1889
Carettochelys insculpta — ERNST & BARBOUR 1989
Carettochelys insculpta — COGGER 2000: 186
Carettochelys insculpta canni WELLS 2002
Carettochelys canni — ARTNER 2003
Carettochelys insculpta — GEORGES & THOMSON 2010 
DistributionPapua New Guinea (Fly, Strickland, Morehead, Lorentz, Stekwa rivers, Lake Jamur),
Australia (Daly, Victoria, Alligator drainages in the Northern Territory)

Type locality: Fly River, Papua New Guinea. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
TypesHolotype: AMS (AM) R3677 
CommentBoulenger (1887) erected the family Carettochelyidae for the genus Carettochelys.

History: Although the volume of the original description was labelled “1887” it was published on 25 May 1886 (M Lambertz, pers. comm., 17 Dec 2012).

Distribution: Map in MACCORD et al. (2003).

Reproduction: C. insculpta has temperature-dependent sex determination (TSD, WEBB et al. 1986, SARRE et al. 2004).

Type species: Carettochelys insculptus RAMSAY 1886 is the type species of the genus Carettochelys RAMSAY 1886: 158 (WERMUTH & MERTENS give 1887 as year). 
  • Barone, S. 2004. The Pig-Nosed Turtle, Carettochelys insculpta, Underwater Glider. Reptilia (GB) (33): 56-60 - get paper here
  • Bonin, F., Devaux, B. & Dupré, A. 2006. Turtles of the World. English translation by P.C.H. Pritchard. Johns Hopkins University Press, 416 pp.
  • Boulenger, G.A. 1887. On a new family of Pleurodiran turtles. Ann. Mag. nat. Hist. (5) 19: 170-172 - get paper here
  • Boulenger, G.A. 1889. Catalogue of the Chelonians, Rhynchocephalians, and Crocodiles in the British Museum (Natural History). British Museum, London, 311 pp.
  • Cann, J. 1972. Notes on some tortoises collected in Northern Australia. Vict. Nat. 89: 165-168
  • Castro, Déborah Praciano de; Maria Juliana Borges-Leite, Daniel Cassiano Lima and Diva Maria Borges- 2013. Parental care in two species of Leptodactylus Fitzinger, 1826 (Anura, Leptodactylidae) in north-eastern Brazil. Herpetology Notes 6: 267-269. - get paper here
  • Cogger, H. G. 1970. First record of the pitted-shelled turtle, Carettochelys insculpta, from Australia. Search 1: 41
  • Cogger,H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Delfino, Massimo; Uwe Fritz and Marcelo R. Sánchez-Villagra 2010. Evolutionary and developmental aspects of phalangeal formula variation in pig-nose and soft-shelled turtles (Carettochelyidae and Trionychidae). Organisms Diversity & Evolution 10 (1): 69-79 - get paper here
  • Doody, J.S., Young, J.E. and Georges, A. 2002. Sex differences in activity and movements in the Pig-Nosed turtle, Carettochelys insculpta, in the wet-dry tropics of Australia. Copeia 2002: 93-103 - get paper here
  • Eisemberg, C. C.; M. Rose, B. Yaru, Y. Amepou and A. Georges 2014. Salinity of the coastal nesting environment and its association with body size in the estuarine pig-nosed turtle Journal of Zoology, DOI: 10.1111/jzo.12179 - get paper here
  • Ernst,C.H. and Barbour,R.W. 1989. Turtles of the World. Smithsonian Institution Press, Washington D.C. - London
  • Georges, A. & Thomson, S. 2010. Diversity of Australasian freshwater turtles, with an annotated synonymy and keys to species. Zootaxa 2496: 1–37 - get paper here
  • Georges, A., Doody, J.S., Eisemberg, C., Alacs, E.A. and Rose, M. 2008. Carettochelys insculpta Ramsay 1886 -- Pig-nosed Turtle, Fly River Turtle. Chelonian Research Monographs 5:9.1-9.17
  • Georges, A.; Alacs, E.; Pauza, M.; Kinginapi, F.; Ona, A. & Eisemberg, C. 2008. Freshwater turtles of the Kikori Drainage, Papua New Guinea, with special reference to the pig-nosed turtle, Carettochelys insculpta. Wildlife Research 35: 700–711
  • Georges, Arthur, Carla Eisemberg, Yolarnie Amepou and Eric Manasi. 2014. Turtle conservation challenges in papua New Guinea. Turtle Survival 2014: 22-24 - get paper here
  • Joyce, Walter G.; James F. Parham , Tyler R. Lyson , Rachel C. M. Warnock and Philip C. J. Donoghue 2013. A Divergence Dating Analysis of Turtles Using Fossil Calibrations: An Example of Best Practices. Journal of Paleontology, 87(4):612-634 - get paper here
  • Kaiser, H.; Crother, B.I.; Kelly, C.M.R.; Luiselli, L.; O’Shea, M.; Ota, H.; Passos, P.; Schleip, W. 2013. Best Practices: In the 21st Century, Taxonomic Decisions in Herpetology are Acceptable Only When Supported by a Body of Evidence and Published via Peer-Review. Herpetological Review 44 (1): 8-23
  • Li, Xiao-san; Nie, Liu-wang; Wang, Ling; Xiong, Lei; Zhou, Ke 2010. The mitochondrial genome complete sequence and organization of the Pig-nosed Turtle Carettochelys insculpta (Testudines, Carettochelyidae) and its phylogeny position in Testudines. Amphibia-Reptilia 31 (4): 541-551 - get paper here
  • Longman, H.A. 1913. Herpetological Notes. Part I. Systematic. Memoirs of the Queensland Museum Brisbane, 2: 39-45
  • Lourenço, João M.; Julien Claude, Nicolas Galtier, Ylenia Chiari 2011. Dating cryptodiran nodes: Origin and diversification of the turtle superfamily Testudinoidea. Molecular Phylogenetics and Evolution 62 (1): - get paper here
  • McCord, W.P.; Joseph-Ouni, M. & Cann, J. 2003. Chelonian Illustrations #7. Short-neck, Western Swamp, and Pig-Nose Turtles from Australia and New Guinea. Reptilia (GB) (27): 64-68 - get paper here
  • Ramsay, E. P. 1886. On a new genus and species of fresh water tortoise from the Fly River, New Guinea. Proc. Linn. Soc., New South Wales (2) 1 (Part 1): 158-162 - get paper here
  • Sarre, S.D.; Georges, A. & Quinn, A. 2004. The ends of a continuum: genetic and temperature-dependent sex determination in reptiles. BioEssays 26: 639–645
  • Schodde, R., Mason, I. & Wolfe, T. O. 1972. Further records of the pitted-shelled turtle (Carettochelys insculpta) from Australia. Trans. R. Soc. S. Aust. 96: 115-117
  • Schultze-Westrum, Thomas 1995. The Papua turtle from New Guinea. Bull. Chicago Herp. Soc. 30 (9): 189-192 - get paper here
  • Siebenrock, F. 1913. Schildkröten aus Syrien und Mesopotamien. Ann. Naturhist. Mus. Wien 27: 171-225 - get paper here
  • Waite, Edgar R. 1905. The osteology of the New Guinea turtle (Carettochelys insculpta, Ramsay). Rec. Austral. Mus. 6 (2): 110-118 - get paper here
  • Webb GJW, Choquenot D, Whitehead P. 1986. Nests, eggs and embryonic development of Carettochelys insculpta (Chelonia: Carettochelidae) from northern Australia. J Zool 1B:521–550.
  • Wells, Richard W. 2002. A New Subspecies of Carettochelys (Reptilia: Carettochelydidae) from Northern Australia - Carettochelys insculpta canni ssp. nov. Australian Biodiversity Record (1):1-7
  • Wilson,S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
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