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Chelonoidis denticulatus (LINNAEUS, 1766)

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Higher TaxaTestudinidae, Testudinoidea, Testudines (turtles) 
Subspecies 
Common NamesE: Yellow-footed Tortoise
G: Waldschildkröte 
SynonymTestudo denticulata LINNAEUS 1766
Testudo tabulata WALBAUM 1782 (nomen illegitimum)
Testudo tabulata SCHOEPFF 1792
Testudo tessellata SCHNEIDER 1792
Testudo hercules SPIX 1824
Testudo sculpta SPIX 1824
Testudo cagado SPIX 1824
Testudo planata GMELIN 1831 (nomen nudum)
Testudo foveolata SCHINZ 1833 (nomen nudum) ?
Testudo tabulata — DUMÉRIL & BIBRON 1835: 89 (?)
Testudo Tabulata — DUMÉRIL & BIBRON 1854: 220
Testudo tabulata — KOSLOWSKY 1898
Geochelone denticulata — WILLIAMS 1960
Geochelone denticulata — DUELLMAN 1978: 194
Chelonoidis denticulata — OBST 1985
Geochelone denticulata — GORZULA & SEÑARIS 1999
Chelonoidis denticulata — MCCORD & JOSEPH-OUNI 2004
Geochelone denticulata — MORAVEC & APARICIO 2005
Chelonoidis denticulata — GERLACH (pers. comm.)
Chelonoidis denticulata — LE et al. 2006
Geochelone denticulata — WHITHWORTH & BEIRNE 2011
Chelonoidis denticulata — MCNISH 2011
Chelonoidis denticulatus — OLSON & DAVID 2014
Chelonoidis denticulata — TTWG 2014: 393
Chelonoidis denticulatus — TTWG 2017: 124 
DistributionSE Venezuela, Caribbean lowlands of Guyana, French Guiana, and Suriname; Brazil (Amazon Basin, isolated range in E Brazil, Espírito Santo, Pará), E Ecuador, Colombia, NE Peru (Pasco), N/E Bolivia, Trinidad, Guadeloupe.

Type locality: in error (see comment)  
Reproductionoviparous. 
TypesHolotype: NRM (NHRM) DeGeer collection 21, according to Andersson 1900. 
Diagnosis 
CommentTerra typica: "Virginia," U.S.A.; in error. LOFTIN (1965) reports this species from Panama but assumes that it most likely was an escapee from a pet shop.

Reaches 70 cm in carapace length (males), although usually most specimens do not grow beyond 50 cm. Males are smaller than females. 
Etymology 
References
  • Andersson, L.G. 1900. Catalogue of Linnean type-specimens of Linnaeus's Reptilia in the Royal Museum in Stockholm. Bihang till Konglika Svenska Vetenskaps-Akademiens. Handlingar. Stockholm. (4) 26 (1): 1-29. - get paper here
  • Barrio-Amorós, César L. & Brewer-Carias, Charles 2008. Herpetological results of the 2002 expedition to Sarisari–ama, a tepui in Venezuelan Guayana, with the description of five new species. Zootaxa 1942: 1-68 - get paper here
  • Bernarde, Paulo Sérgio;, Reginaldo Assêncio Machado & Luiz Carlos Batista Turci 2011. Herpetofauna da área do Igarapé Esperança na Reserva Extrativista Riozinho da Liberdade, Acre – Brasil. Biota Neotrop. 11 (3): 117-144 - get paper here
  • Bonin, F., Devaux, B. & Dupré, A. 2006. Turtles of the World. English translation by P.C.H. Pritchard. Johns Hopkins University Press, 416 pp.
  • Castaño-Mora, O.V., and M. Lugo R. 1981. Estudio comparativo del comportamiento de dos especies de morrocoy: Geochelone carbonaria y Geochelone denticulata y aspectos comparables de su morfología esterna. Cespedesia 10:55-122
  • CATENAZZI, A., LEHR, E. & VON MAY, R. 2013. The amphibians and reptiles of Manu National Park and its buffer zone, Amazon basin and eastern slopes of the Andes, Peru. Biota Neotrop. 13(4): 269-283
  • Cole, Charles J.; Carol R. Townsend, Robert P. Reynolds, Ross D. MacCulloch, and Amy Lathrop 2013. Amphibians and reptiles of Guyana, South America: illustrated keys, annotated species accounts, and a biogeographic synopsis. Proceedings of the Biological Society of Washington 125 (4): 317-578; plates: 580-620 - get paper here
  • Duellman, W. E. 1978. The biology of an equatorial herpetofauna in Amazonian Ecuador. Misc. Publ. Univ. Kans. Mus. Nat. Hist. 65: 1-352 - get paper here
  • Duellman, W. E. 2005. Cusco Amazónico: The Lives of Amphibians and Reptiles in an Amazonian Rainforest. Comstock Pub Assoc.
  • Duellman, W.E., & Salas, A.W. 1991. Annotated checklist of the amphibians and reptiles of Cuzco Amazonico, Peru. Occas. Papers Mus. of Natur. Hist., Univ. of Kansas, Lawrence (143): 13 pp. - get paper here
  • Duméril, A. M.C., G. BIBRON & A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 9. Paris, XX + 440 S. - get paper here
  • Duméril, A.M.C., and G. Bibron. 1835. Erpétologie Générale ou Histoire Naturelle Complète des Reptiles, Vol. 2. Librairie Encyclopédique de Roret, Paris, iv + 680 p. - get paper here
  • Ernst,C.H. and Barbour,R.W. 1989. Turtles of the World. Smithsonian Institution Press, Washington D.C. - London
  • Ferronato, B. et al. 2011. New locality records for chelonians (Testudines: Chelidae, Podocnemididae, Testudinidae) from Departamento de Pasco, Peru. Herpetology Notes 4: 219-224 - get paper here
  • Gaffney, E. S. 1979. Comparative cranial morphology of Recent and fossil turtles. Bull. Amer. Mus. Nat. Hist. 164 (2): 67-376
  • Gorzula, Stefan & Senaris, J. Celsa 1999. In: Contribution to the herpetofauna of the Venezuelan Guayana. I: a data base. Scientia Guaianae, Caracas, No. 8 [1998], 269+ pp.; ISBN 980-6020-48-0
  • Gray,J. E. 1866. Notes on some young specimens of tortoises (Testudo). Proc. Zool. Soc. London 1866: 305-306 - get paper here
  • Hoogmoed,M.S. and Gruber,U. 1983. Spix and Wagler type specimens of reptiles and amphibians in the Natural History Musea in Munich (Germany) and Leiden (The Netherlands). Spixiana Suppl. 9: 319-415 - get paper here
  • Humair, Robert. 2013. Scutellation anomalies in a Yellow-footed Tortoise, Chelonoidis denticulata. Radiata 22 (4): 35
  • Humair, Robert. 2013. Dimensions and weights of South American Yellow-footed Tortoises, Chelonoidis denticulata (Linnaeus, 1766), in nature and human care. Radiata 22 (4): 21-30
  • Knaack, J. 2006. Beobachtungen zum Vorkommen von Geochelone (Chelonoidis) im südlichen Bolivien. Reptilia (Münster) 11 (59): 77-82 - get paper here
  • Le, Minh; Christopher J. Raxworthy, William P. McCord and Lisa Mertz 2006. A molecular phylogeny of tortoises (Testudines: Testudinidae) based on mitochondrial and nuclear genes. Molecular Phylogenetics and Evolution 450 (2): 517-531 - get paper here
  • Lehr, E. 2001. New records for amphibians and reptiles from Departamentos Pasco and Ucayali, Peru. Herpetological Review 32 (2): 130-132 - get paper here
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  • Lodge, T. M., T. F. Elliott and T. W. Henkel 2016. Chelonoidis denticulata (Yellow-footed Tortoise) diet. Herpetological Review 47(3): 443.
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  • McCord, W.P. & Joseph-Ouni, M. 2004. Chelonian Illustrations #14: North and South American Tortoises. Reptilia (GB) (34): 47-50 - get paper here
  • McNish, T. 2011. La fauna del archipiélago de San Andrés, Providencia y Santa Catalina, Colombia, Sudamérica. Colomba Andina de Impresos, ISBN 978-958-99518-1-1
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  • Morais, D. H., et al. 2017. Patterns of parasitism by Amblyomma humerale (Acari: Ixodidae) on Chelonoidis denticulata (Testudines: Testudinidae) in the Brazilian midwest. Herpetological Review 48 (2): 316-319
  • Morato, S.A.A.; Calixto, P.O.; Mendes, L.R.; Gomes, R.; Galatti, U.; Trein, F.L.; Oliveira, F.S.; Ferreira, G.N. 2014. Guia fotográfico de identificação da herpetofauna da Floresta Nacional de Saracá-Taquera, Estado do Pará. Curitiba: STCP Engenharia de Projetos Ltda.; Porto Trombetas: MRN – Mineração Rio do Norte S.A.; 213 p. - get paper here
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