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Cnemaspis chanardi GRISMER, SUMONTHA, COTA, GRISMER, WOOD, PAUWELS & KUNYA, 2010

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Chan-ard’s Rock Gecko
Thai: Djing Djok Niew Yaow Tanya 
SynonymCnemaspis chanardi GRISMER, SUMONTHA, COTA, GRISMER, WOOD, PAUWELS & KUNYA 2010
Gonatodes siamensis SMITH 1930:16
Cnemaspis siamensis SMITH 1935:71
Cnemaspis siamensis — TAYLOR 1963: 740 (part.)
Cnemaspis chanardi — GRISMER et al. 2014: 52
Cnemaspis chanardi — COTA et al. 2022 
DistributionS Thailand (foothills of the Nakhon Si Thammarat and Sankalakhiri Mountains and lowland areas to the west, extending from the southern terminus of the Isthmus of Kra in Donsak District, Surat Thani Province, southward along the western foothills through Khao Chong and Nayong district, Trang Province to at least the Phuphaphet Cave, Satun Province in the south. It probably continues approximately 45 km further south to the Banjaran Nakawan mountains that form the physiographic barrier between Thailand and Peninsular Malaysia. It extends westward from the foothills through the lowlands to at least Khlong Thom District, Krabi Province. It ranges from near sea level at Khlong Thom to just under 600 m at Khao Chong. Smith (1930) reported specimens from Khao Whip (=Khao Wang Hip), Nakhon Si Thammarat Province but the elevation is unknown. Its presence on Ko Tao Island, Surat Thani Province, approximately 85 km off the coast from Muang District, Chumpon Province, as opposed to the presence of Cnemaspis siamensis which is geographically closer to Ko Tao Island, is consistent with the geological history of this part of Peninsular Thailand. The island chain consisting of Ko Tao and the intervening islands Ko Samui and Ko Phangan, are offshore extensions of the Nakhon Si Thammarat Mountains to which they were connected during the last glacial maximum (Sathiamurthy & Voris 2006) and lie to the east of the Isthmus of Kra. It is likely that C. chanardi also occurs on Ko Samui and Ko Phangan; see also map [Fig. 1] in GRISMER et al. 2010)

Type locality: at Ban Chong, Chong, Nayong District, Trang Province, Thailand.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: THNHM 6983, adult male, collected on 22 June 2005. 
DiagnosisDiagnosis: Adult males reaching 40.1 mm SVL, adult females reaching 38.7 mm SVL; 7–10 supralabials; 6–8 infralabials; gulars smooth; forearm, subtibials, ventrals, subcaudals, and dorsal tubercles keeled; 20–30 paravertebral tubercles; tubercles on flanks not linearly arranged; ventrolateral caudal tubercles absent; caudal tubercles do not encircle tail; no lateral, caudal tubercles within lateral, caudal furrow; median row of subcaudals keeled, slightly enlarged; 6–8 precloacal, pore-bearing scales in adult males separated medially by non-pore-bearing scales; pores round; one postcloacal tubercle; shield-like subtibials and enlarged, submetatarsals absent; 25–30 subdigital lamellae on fourth toe; no dark, longitudinal gular markings or blotches; head not yellow in adult males; no dark patch on shoulder or neck enclosing a white to yellow ocellus; yellow to white, prescapular crescent present. These differences are summarized across all species in TABLES 1 and 2 in GRISMER et al. 2010, and Grismer et al. 2014 (Table 6). 
CommentThis species was previously considered as C. siamensis. Cnemaspis chanardi is most similar to C. kamolnorranathi, C. roticanai, C. siamensis, and C. vandeventeri of Peninsular Thailand.

Distribution: See map in Grismer 2020: 548 (Fig. 1), Grismer et al. 2014: 17 (Fig. 3). 
Etymology“The specific epithet chanardi, a masculine name in the genitive case, is in reference to Mr. Tanya Chan-ard of the Thailand Natural History Museum, National Science Museum, Bangkok for his extensive contributions to the herpetology of Thailand, his gracious assistance with this project, and for collecting much of the material used in this study” [from GRISMER et al. 2010]. 
References
  • Ampai N, Rujirawan A, Yodthong S, Termprayoon K, Stuart BL, Wood Jr PL, Aowphol A 2022. Hidden diversity of rock geckos within the Cnemaspis siamensis species group (Gekkonidae, Squamata): genetic and morphological data from southern Thailand reveal two new insular species and verify the phylogenetic affinities of C. chanardi and C. kam ZooKeys 1125: 115–158 - get paper here
  • Ampai N, Wood Jr PL, Stuart BL, Aowphol A 2020. Integrative taxonomy of the rock-dwelling gecko Cnemaspis siamensis complex (Squamata, Gekkonidae) reveals a new species from Nakhon Si Thammarat Province, southern Thailand. ZooKeys 932: 129-159 - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Cota, M., Makchai, S., & Pongcharoen, C. 2022. Updated Catalogue of the Amphibian and Reptile Type Specimens of the Natural History Museum, National Science Museum, Thailand. Thai Specimens 21: 311––5407
  • GRISMER, L. LEE; MONTRI SUMONTHA, MICHAEL COTA, JESSE L. GRISMER, PERRY L. WOOD, JR., OLIVIER S. G. PAUWELS & KIRATI KUNYA 2010. A revision and redescription of the rock gecko Cnemaspis siamensis (Taylor 1925) (Squamata: Gekkonidae) from Peninsular Thailand with descriptions of seven new species. Zootaxa 2576: 1–55 - get paper here
  • GRISMER, L. LEE; PLATON V. YUSHCHENKO, PARINYA PAWANGKHANANT, ROMAN A. NAZAROV, MALI NAIDUANGCHAN, CHATMONGKON SUWANNAPOOM, NIKOLAY A. POYARKOV 2020. A new species of Cnemaspis Strauch (Squamata: Gekkonidae) of the C. siamensis group from Tenasserim Mountains, Thailand. Zootaxa 4852 (5): 547–564; erratum in Zootaxa 4951 (3): 599 (DOI: 10.11646/zootaxa.4951.3.11) - get paper here
  • Grismer, L.L.; PERRY JR L. WOOD, SHAHRUL ANUAR, AWAL RIYANTO, NORHAYATI AHMAD, MOHD A. MUIN, MONTRI SUMONTHA, JESSE L. GRISMER, CHAN KIN ONN, EVAN S. H. QUAH, OLIVIER S. A. PAUWELS 2014. Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia. Zootaxa 3880 (1): 001–147 - get paper here
  • Nguyen, H.N., Hung, C., Yang, M. et al. 2020. Sympatric competitors have driven the evolution of temporal activity patterns in Cnemaspis geckos in Southeast Asia. Sci Rep 10: 27 - get paper here
  • Rujirawan, A., Yodthong, S., Ampai, N., Termprayoon, K., Aksornneam, A., Stuart, B. L., & Aowphol, A. 2022. A new rock gecko in the Cnemaspis siamensis group (Reptilia, Gekkonidae) from Kanchanaburi Province, western Thailand. Zoosystematics and Evolution, 98(2), 345-363 - get paper here
  • WOOD, PERRY JR. L.; EVAN S.H. QUAH, SHAHRUL ANUAR M.S., MOHD ABDUL MUIN 2013. A new species of lowland karst dwelling Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from northwestern Peninsular Malaysia. Zootaxa 3691 (5): 538–558 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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