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Cnemaspis affinis (STOLICZKA, 1870)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Stoliczka's Gecko, Pinang Island Rock Gecko
G: Penang Dünnfingergecko 
SynonymCyrtodactylus affinis STOLICZKA 1870: 167
Cyrtodactylus affinis STOLICZKA 1870: 107
Gymnodactylus affinis — BOULENGER 1885: 42
Gonatodes penangensis FLOWER 1897 (fide TAYLOR 1963)
Gonatodes affinis — DE ROOIJ 1915: 24
Cnemaspis affinis — TAYLOR 1963: 744
Gonatodes affinis — FLOWER 1898
Cnemaspis affinis — GRANDISON 1972: 80
Cnemaspis affinis — KLUGE 1993
Cnemaspis (Cnemaspis) affinis — RÖSLER 2000: 62
Cnemaspis affinis — GRISMER 2011
Cnemaspis affinis — GRISMER et al. 2014: 66 
DistributionPeninular Malaysia (Pulau Pinang)

Type locality: Penang Hills.  
Reproductionoviparous 
TypesHolotype: ZSI 5964 
DiagnosisDiagnosis: Maximum SVL 50.8 mm; 9–13 supralabials; 8–10 infralabials; ventral scales keeled; five or six discontinuous, pore-bearing precloacal scales with round pores; 20–28 paravertebral tubercles; dorsal tubercles not linearly arranged, present on flanks; tubercles absent from lateral caudal furrows; ventrolateral caudal tubercles absent; lateral caudal row present; caudal tubercles not encircling tail; all subcaudals keeled, no enlarged median scale row; two postcloacal tubercles on each side of tail base; no enlarged femoral or subtibial scales; subtibials keeled; no submetatarsal scales on first toe; 28 or 29 subdigital fourth toe lamellae; gular region yellow in males; single ocellus in shoulder region in males, yellow post-scapular band in males; and transverse yellow bars on flanks.These differences are summarized across all Southeast Asian species in Grismer et al. (2008b: Table 1) and GRISMER et al. 2014 (Tables 6,7). 
CommentDistribution: Das & Bauer (1998) and GRISMER & ONN (2009) couldn’t find any evidence for Cnemaspis affinis on Borneo and thus Indonesia. Grismer (2010) says this species is endemic to Pulau Pinang. Reports from Thailand and Malacca probably represent other species. 
References
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Das I 1994. Cnemaspis gordongekkoi, a new gecko from Lombok, Indonesia, and the biogeography of oriental species of Cnemaspis (Squamata: Sauria: Gekkonidae). Hamadryad 18: 1-9 [1993] - get paper here
  • Das, I. 2005. Revision of the Genus Cnemaspis Strauch, 1887 (Sauria: Gekkonidae), from the Mentawai and Adjacent Archipelagos off Western Sumatra, Indonesia, with the Description of Four New Species. Journal of Herpetology 39 (2): 233–247 - get paper here
  • Das, I. & Grismer, L.L. 2003. Two new species of Cnemaspis Strauch 1887 (Squamata, Gekkonidae) from the Seribuat Archipelago, Pahang and Johor States, West Malaysia. Herpetologica 59 (4): 544-552 - get paper here
  • Das, I. & Yaakob, N. 2007. Status of knowledge of the Malaysian herpetofauna. In Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. in: L. S. L. Chua, L. G. Kirton & L. G. Saw (eds.), Status of biological diversity in Malaysia & threat assessment of plant species in Malaysia. Forest Research Institute Malaysia, Kepong, pp. 31-81
  • Das,I. & A. M. BAUER 1998. Systematics and biogeography of Bornean geckos of the genus Cnemaspis Strauch, 1887 (Sauria: Gekkonidae), with the description of a new species. Raffles Bull. Zool. 46 (1): 11-28. - get paper here
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Flower, S. S. 1897. Notes on a collection of reptiles and batrachians made in the Malay Peninsula in 1895-96; With a list of the species recorded in that region. Proc. Zool. Soc. London 1896: 856—914. - get paper here
  • Grandison, A.G.C. 1972. The Gunong Benom Expedition 1967. 5. Reptiles and amphibians of Gunong Benom with a description of a new species of Macrocalamus. Bull. Br. Mus. nat. Hist. (Zool.), London, 23: 45-101. - get paper here
  • Grismer, J.L.; Grismer, L.L. & Chav T. 2010. New Species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Southwestern Cambodia. Journal of Herpetology 44 (1): 28–36 - get paper here
  • Grismer, L. Lee & Chan K. Onn 2008. A new species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. Zootaxa 1771: 1–15 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L. Lee; Chan K. Onn; Nasir, N. & Sumontha, M. 2008. A new species of karst dwelling gecko (genus Cnemaspis Strauch 1887) from the border region of Thailand and Peninsular Malaysia. Zootaxa 1875: 51–68 - get paper here
  • GRISMER, L. LEE; MONTRI SUMONTHA, MICHAEL COTA, JESSE L. GRISMER, PERRY L. WOOD, JR., OLIVIER S. G. PAUWELS & KIRATI KUNYA 2010. A revision and redescription of the rock gecko Cnemaspis siamensis (Taylor 1925) (Squamata: Gekkonidae) from Peninsular Thailand with descriptions of seven new species. Zootaxa 2576: 1–55 - get paper here
  • GRISMER, L. LEE; NORHAYATI AHMAD, CHAN KIN ONN, DAICUS BELABUT, MUIN, M. A., PERRY L. WOOD, JR, & JESSE L. GRISMER 2009. Two new diminutive species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Peninsular Malaysia. Zootaxa 2019: 40-56 - get paper here
  • GRISMER, L. LEE; P. L. WOOD, JR., EVAN S. H. QUAH, SHAHRUL ANUAR, EHWAN NGADI & NORHAYATI AHMAD 2015. A new insular species of Rock Gecko (Cnemaspis Boulenger) from Pulau Langkawi, Kedah, Peninsular Malaysia. Zootaxa 3985 (2): 203–218 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Grismer, L.L. & Das, I. 2006. A new species of gekkonid lizard of the genus Cnemaspis STRAUCH 1887 from Pulau Pemanggil, Johor, West Malaysia. Herpetological Natural History 10 (1): 1-7 - get paper here
  • Grismer, L.L. & Ngo, V.T. 2007. Four new species of the gekkonid genus Cnemaspis Strauch 1887 (Reptilia: Squamata) from Southern Vietnam. Herpetologica 63 (4): 482-500 - get paper here
  • Grismer, L.L.; Grismer, J.L.; Wood Jr., P.L.; Chan K. Onn 2008. The distribution, taxonomy, and redescription of the geckos Cnemaspis affinis (Stoliczka 1887) and C. flavolineata (Nicholls 1949) with descriptions of a new montane species and two new lowland, karst- dwelling species from Peninsular Malaysia. Zootaxa 1931: 1–24 - get paper here
  • Grismer, L.L.; PERRY JR L. WOOD, SHAHRUL ANUAR, AWAL RIYANTO, NORHAYATI AHMAD, MOHD A. MUIN, MONTRI SUMONTHA, JESSE L. GRISMER, CHAN KIN ONN, EVAN S. H. QUAH, OLIVIER S. A. PAUWELS 2014. Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia. Zootaxa 3880 (1): 001–147 - get paper here
  • Grismer, L.L.; PERRY L. WOOD JR., MAKETAB MOHAMED, KIN ONN CHAN, HEATHER M. HEINZ, ALEX S-I. SUMARLI, JACOB A. CHAN, ARIEL I. LOREDO 2013. A new species of karst-adapted Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) from a threatened karst region in Pahang, Peninsular Malaysia. Zootaxa 3746 (3): 463–472 - get paper here
  • Nguyen, H.N., Hung, C., Yang, M. et al. 2020. Sympatric competitors have driven the evolution of temporal activity patterns in Cnemaspis geckos in Southeast Asia. Sci Rep 10: 27 - get paper here
  • Onn, Chan Kin; L. L. Grismer, Shahrul Anuar, E. Quah, M.A. Muin, A.E. Savage, J.L. Grismer, Norhayati Ahmad, A.C. Remigio, L.F. Greer 2010. A new endemic rock Gecko Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia. Zootaxa 2576: 59–68 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Stoliczka, F. 1870. Observations on some Indian and Malayan Amphibia and Reptilia. J. Asiat. Soc. Bengal 39: 134-228. - get paper here
  • Stoliczka, F. 1870. Observations of some Indian and Malayan Amphibia and Reptilia (abstract). Ann. Mag. Nat. Hist. (4) 6: 105-109 - get paper here
  • Taylor, E.H. 1963. The lizards of Thailand. Univ. Kansas Sci. Bull. 44: 687-1077. - get paper here
  • WOOD, PERRY JR. L.; EVAN S.H. QUAH, SHAHRUL ANUAR M.S., MOHD ABDUL MUIN 2013. A new species of lowland karst dwelling Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from northwestern Peninsular Malaysia. Zootaxa 3691 (5): 538–558 - get paper here
 
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