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Cnemaspis bidongensis GRISMER, WOOD, AHMAD, SUMARLI, VAZQUEZ, ISMAIL, NANCE, MOHD-AMIN, OTHMAN, RIZAIJESSIKA, KUSS, MURDOCH & COBOS, 2014

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Pulau Bidong Rock Gecko
Malay: Cicak Batu Pulau Bidong 
SynonymCnemaspis bidongensis GRISMER, WOOD, AHMAD, SUMARLI, VAZQUEZ, ISMAIL, NANCE, MOHD-AMIN, OTHMAN, RIZAIJESSIKA, KUSS, MURDOCH & COBOS 2014
Cnemaspis bidongensis — GRISMER et al. 2014: 124 
DistributionPeninsular Malaysia (Pulau Bidong)

Type locality: Pulau Bidong, Terengganu, Peninsular Malaysia (5°37.201’ N 103°03.244’ E; 49 m elevation  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: LSUHC 11455, adult female, collected on 26 August 2013 by Jacob A. Chan at 2200 hrs. Paratype. Adult males (LSUHC 11447, 11452–54) and adult female LSUHC 11451 bear the same data as the holotype except they were collected between 1800 and 2400 hrs. 
DiagnosisDiagnosis: Cnemaspis bidongensis sp. nov. differs from all other Southeast Asian species of Cnemaspis in having the unique combination of adult males reaching at least 56.1 mm SVL, adult females reaching 58.1 mm SVL; nine or 10 supralabials; 7–9 infralabials; enlarged, elongate mental scale; keeled ventrals; no precloacal pores; moderately prominent, randomly arranged, dorsal tubercles; 21–26 paravertebral tubercles; no tubercles on lower flanks caudal tubercles encircling tail; no tubercles in lateral caudal furrows; ventrolateral caudal tubercles present anteriorly; subcaudals keeled; a median row of enlarged, keeled subcaudals; one or two postcloacal tubercles on each side; no enlarged femoral, subtibial, or submetatarsal scales; subtibials keeled; 26–30 subdigital lamellae on fourth toe; dark and light caudal bands distinct in both sexes; and no dark mottling on lateral portions of belly. These differences are summarized across all species of the C. kendallii species group in Table 1 and all across Southeast Asian species in Wood et al. (2013:Table 4) and GRISMER et al. 2014 (Tables 6,7). 
CommentAbundance: only known from its original description (Meiri et al. 2017).

Distribution: See map in Grismer et al. 2014: 18 (Fig. 4). 
EtymologyNamed after the type locality. 
References
  • Goldberg, S. R., C. R. Bursey and L. L. Grismer 2016. Cnemaspis bidongensis (Pulau Bidong Rock Gecko) endoparasites. Herpetological Review 47(3): 462. - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L.; PERRY JR L. WOOD, SHAHRUL ANUAR, AWAL RIYANTO, NORHAYATI AHMAD, MOHD A. MUIN, MONTRI SUMONTHA, JESSE L. GRISMER, CHAN KIN ONN, EVAN S. H. QUAH, OLIVIER S. A. PAUWELS 2014. Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia. Zootaxa 3880 (1): 001–147 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR., AMIRRUDIN B. AHMAD, ALEXANDRA S.-I. SUMARLI, JESSIKA J. VAZQUEZ, LUKMAN H. B. ISMAIL, RONALD NANCE, MUHAMMAD AFIF B. MOHD-AMIN, MOHAMAD N. A. B. OTHMAN, SYED A. RIZAIJESSIKA, MARIA KUSS, MATTHEW MURDOCH, ANTHONY COB 2014. A new species of insular Rock Gecko (Genus Cnemaspis Strauch, 1887) from the Bidong Archipelago, Terengganu, Peninsular Malaysia. Zootaxa 3755 (5): 447–456 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Nguyen, H.N., Hung, C., Yang, M. et al. 2020. Sympatric competitors have driven the evolution of temporal activity patterns in Cnemaspis geckos in Southeast Asia. Sci Rep 10: 27 - get paper here
  • RIYANTO, AWAL; MISBAHUL MUNIR, ANDRI I. S. MARTAMENGGALA, YULI SULISTYA FITRIANA, AMIR HAMIDY 2019. Hiding in plain sight on Gunung Muria: A new species and first record of rock gecko (Cnemaspis Strauch, 1887; Squamata, Gekkonidae) from Java, Indonesia. Zootaxa 4608 (1): 155–173 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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