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Cnemaspis kivulegedarai KARUNARATHNA, POYARKOV, DE SILVA, MADAWALA, BOTEJUE, GORIN, SURASINGHE, GABADAGE, UKUWELA & BAUER, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common Names 
SynonymCnemaspis kivulegedarai KARUNARATHNA, POYARKOV, DE SILVA, MADAWALA, BOTEJUE, GORIN, SURASINGHE, GABADAGE, UKUWELA & BAUER 2019 
DistributionSri Lanka (Central Province)

Type locality: granite wall in Keerthibandarapura, Walapane, Nuwara Eliya District, Central Province, Sri Lanka (7.127933° N, 80.867364° E, WGS 1984; elevation 539 m  
Reproduction 
TypesHolotype. NMSL.2019.08.01, adult male, 28.5 mm SVL (Fig. 19),
collected around 14.00 hrs) on 17 June 2018 by Suranjan Karunarathna and Anslem de Silva.
Paratypes. NMSL.2019.08.02, adult female, 31.2 mm SVL, and
NMSL.2019.08.03, adult female, 29.8 mm SVL, collected from a
large granite wall in Keerthibandarapura, Walapane, Nuwara Eliya District, Central Province, Sri Lanka (7.140061° N, 80.872467° E, WGS 1984; elevation 562 m; around 15.00 hrs) on 17 June 2018 by Suranjan Karunarathna and Anslem de Silva. 
DiagnosisDiagnosis. Cnemaspis kivulegedarai sp. nov., can be readily distinguished from its Sri Lankan congeners by a combination of the following morphological and meristic characteristics: maximum SVL 31.2 mm; dorsum with heterogeneous, smooth, granular scales; 1 internasal, 2/2 supranasals and 1/1 postnasal present; 30 – 34 interorbital scales; 15 – 17 supraciliaries, 12 canthal scales, 17 – 21 eye to tympanum scales; 3 enlarged postmentals; postmentals bounded by 5 chin scales; chin with smooth granules; gular, pectoral and abdominal scales smooth, subimbricate; 19 belly scales across the venter; 4 – 5 weakly developed tubercles on posterior flank; 131 – 133 paravertebral granules linearly arranged; 2 precloacal pores, 4 – 5 femoral pores on each side in males separated by 9 – 10 unpored anterior femoral scales, 2 unpored posterior femoral scales; 109 – 114 ventral scales; 69 – 76 midbody scales; subcaudals smooth, diamond shaped, in irregular series forming a narrow median row; 7 supralabials; 6 – 7 infralabials; 14 total lamellae on fourth digit of manus, and 15 total lamellae on fourth digit of pes.

Comparisons with other species. Among species of the C. kandiana clade sensu Agarwal et al. (2017) C. kivulegedarai sp. nov. differs from C. amith, C. gotaimbarai sp. nov., C. kumarasinghei, C. latha and C. nandimithrai sp. nov. by having heterogeneous (versus homogeneous) dorsal scales. It also diagnosed from C. pava, C. pulchra, C. samanalensis, C. silvula, C. tropidogaster and C. upendrai by having smooth (versus keeled) pectoral and ventral scales, from C. butewai sp. nov., C. kandiana, C. menikay and C. retigalensis by having smooth (versus keeled) chin and gular scales; from C. kallima by having fewer ventral scales (109 – 114 versus 131 – 138), and fewer total lamellae on digit IV of pes (15 versus 18 – 20); and from C. ingerorum by having more ventral scales (109 – 114 versus 88 – 95), and fewer total lamellae on digit IV of pes (15 versus 17 – 18). The new species, C. kivulegedarai sp. nov. also clearly differs from the species of the C. podihuna clade sensu Agarwal et al. (2017). From C. alwisi, C. gemunu, C. godagedarai, C. hitihami sp. nov., C. kohukumburai sp. nov., C. nilgala, C. phillipsi, C. punctata, C. rajakarunai, C. rammalensis, and C. scalpensis it differs by the presence (versus absence) of precloacal pores and also by the absence of clearly enlarged, hexagonal or subhexagonal subcaudal scales. From C. kandambyi, C. molligodai and C. podihuna it differs by having heterogeneous (versus homogeneous) dorsal scales and by the absence of clearly enlarged, hexagonal or subhexagonalshaped subcaudal scales. 
Comment 
EtymologyThe specific epithet is an eponym Latinized (kivulegedarai) in the masculine genitive singular, honouring ‘‘Kivulegedara Mohottala’ (a warrior) for his valiant feats in the Great Rebellion of 1817 – 1818, which was initiated in Uva-Wellassa. 
References
  • Karunarathna, Suranjan; Nikolay A. Poyarkov, Anslem de Silva, Majintha Madawala, Madhava Botejue, Vladislav A. Gorin, Thilina Surasinghe, Dinesh Gabadage, Kanishka D.B. Ukuwela & Aaron M. Bauer 2019. Integrative taxonomy reveals six new species of day geckos of the genus Cnemaspis Strauch, 1887 (Reptilia: Squamata: Gekkonidae) from geographically-isolated hill forests in Sri Lanka. Vertebrate Zoology 69 (3): 247–298 - get paper here
 
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