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Cnemaspis niyomwanae GRISMER, SUMONTHA, COTA, GRISMER, WOOD, PAUWELS & KUNYA, 2010

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Niyomwan’s Rock Gecko
Thai: Djing Djok Niew Yaow Niyomwan 
SynonymCnemaspis niyomwanae GRISMER, SUMONTHA, COTA, GRISMER, WOOD, PAUWELS & KUNYA 2010
Cnemaspis niyomwanae — GRISMER et al. 2014: 36 
DistributionNE Thailand (border regions of Trang and Satun provinces), W Cambodia

Type locality: Thum Khao Ting, Palean District, Trang Province, Thailand (07°09.943N 99°48.142E) at 28 m in elevation. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproduction 
TypesHolotype: THNHM 15910, adult female. Collected by M. Sumontha on 26 December 2009.
Paratypes. Adult male (ZMKU Rep-000315), adult female (PSUZC-RT 2010.56), subadult males (CUMZ R-2009,6,24-10, KZM 008) from Baan Nam Pud, La-ngu District, Satun Province (07°05.688N 99° 54.732E) at 38–46 m in elevation. ZMKU Rep-000315, CUMZ R-2009,6,24-10 and KZM 008 were collected by Siriwat Dangsri on 6 and 7 October 2009. PSUZC-RT 2010.56 was collected by Thanin Kaewmanee on 24 December 2009. 
CommentComparisons. Cnemaspis niyomwanae sp. nov. can be diagnosed from all other Southeast Asian Cnemaspis in having males bearing alternating red-orange and yellow bands on the forelimbs and enlarged, white, isolated tubercles on the sides of the neck, shoulder and flanks. Its lack of white ocelli in a black shoulder or neck patch separates it from C. affinis, C. biocellata, C. kumpoli, C. mcguirei, C. pseudomcguirei, C. puntatonucalis sp. nov. Its presence of precloacal pores in males further separates it from all other species except C. affinis, C. argus, C. bayuensis, C. biocellata, C. caudanivea, C. chanthaburiensis, C. dringi, C. flavigaster, C. flavolineata, C. karsticola, C. kumpoli, C. mcguirei, C. monachorum, C. nigridia, C. nuicamensis, C. paripari, C. perhentianensis, C. pseudomcguirei, C. chanardi sp. nov., C. vandeventeri sp. nov., C. kamolnorranathi sp. nov., C. narathiwatensis sp. nov., C. huaseesom sp. nov., and C. roticanai.

Diagnosis. Adult males reaching 45.9 mm SVL, adult females reaching 56.8 mm SVL; 8–11 supralabials; 6–8 infralabials; gulars smooth; forearm scales keeled; subtibials, ventrals, subcaudals smooth; dorsal tubercles keeled; 26–31 paravertebral tubercles; enlarged, elongate, laterally compressed, white, isolated tubercles on sides of neck, shoulders, and flanks; dorsolateral, lateral, and ventrolateral caudal tubercles usually present only anteriorly; caudal tubercles do not encircle tail; caudal tubercles absent from lateral, caudal furrow; middorsal caudal furrow absent; median row of smooth, enlarged subcaudals; three pore- bearing, precloacal scales in males; one or two postcloacal tubercles; shield-like subtibials and enlarged, submetatarsals absent; 31–34 subdigital lamellae on fourth toe; yellow and orange alternating bands on forearms and forelimbs in subadult and adult males; paired, yellow, rostral stripes in subadult and adult males; paried, cream to white, circular, occipital blotches; large, cream to white, paravertebral butterfly-shaped dorsal blotches; no dark, longitudinal, gular markings or blotches; head not yellow in adult males; no black neck patch enclosing a white to yellow ocellus; no dark shoulder patch enclosing white to yellow ocellus; no prominent, yellow to white, prescapular crescent or transverse bars on flanks. These differences are summarized across all species in TABLES 1 and 2 in GRISMER et al. 2010. 
EtymologyThis species is named in honor of Ms. Piyawan Niyomwan, Thai herpetologist who has worked for many years making significant contributions to our knowledge of the distribution of the amphibians and reptiles of Thailand. 
References
  • GRISMER, L. LEE; MONTRI SUMONTHA, MICHAEL COTA, JESSE L. GRISMER, PERRY L. WOOD, JR., OLIVIER S. G. PAUWELS & KIRATI KUNYA 2010. A revision and redescription of the rock gecko Cnemaspis siamensis (Taylor 1925) (Squamata: Gekkonidae) from Peninsular Thailand with descriptions of seven new species. Zootaxa 2576: 1–55 - get paper here
  • GRISMER, L. LEE; PERRY JR L. WOOD, SHAHRUL ANUAR, AWAL RIYANTO, NORHAYATI AHMAD, MOHD A. MUIN, MONTRI SUMONTHA, JESSE L. GRISMER, CHAN KIN ONN, EVAN S. H. QUAH, OLIVIER S. A. PAUWELS 2014. Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia. Zootaxa 3880 (1): 001–147 - get paper here
  • WOOD, PERRY JR. L.; EVAN S.H. QUAH, SHAHRUL ANUAR M.S., MOHD ABDUL MUIN 2013. A new species of lowland karst dwelling Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from northwestern Peninsular Malaysia. Zootaxa 3691 (5): 538–558 - get paper here
 
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