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Cnemaspis shahruli GRISMER, CHAN, QUAH, MUIN, SAVAGE, GRISMER, AHMAD, GREER & REMEGIO, 2010

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Shahrul’s Rock Gecko 
SynonymCnemaspis shahruli GRISMER, CHAN, QUAH, MUIN, SAVAGE, GRISMER, AHMAD, GREER & REMEGIO 2010
Cnemaspis shahruli — GRISMER 2011
Cnemaspis shahruli — GRISMER et al. 2014: 72 
DistributionW Malaysia (Penang and Jerejak Islands, Penang; Sungai Sedim, Kedah; Pulau Pangkor, Perak)

Type locality: Telok Bahang Recreational Forest Reserve, Penang (05°27.233’N 100°12.324’E), 67 m elevation.  
Reproductionoviparous (manual and phylogenetic imputation, fide Zimin et al. 2022) 
TypesHolotype: ZRC 2.6898; adult male, collected on 14 March 2010 by Evan Quah. Paratypes. ZRC 
DiagnosisDiagnosis: Maximum SVL 36.5 mm; 10 or 11 supralabials; 8–10 infralabials; ventral scales keeled; no precloacal pores; 19–23 paravertebral tubercles; body tubercles randomly arranged, present on flanks; tubercles within lateral caudal furrows; ventrolateral caudal tubercles absent anteriorly; lateral row of caudal tubercles present anteriorly; caudal tubercles not encircling tail; all subcaudals keeled, no enlarged median row; 1–3 postcloacal tubercles on each side of tail base; no enlarged femoral or subtibial scales; subtibials keeled; usually no enlarged submetatarsal scales on first toe; 21–30 subdigital fourth toe lamellae; light-colored vertebral stripe variably present; gular region, throat and pectoral region yellow in males; dark, central, elongate marking occurs in mental region; a single ocellus in the shoulder region in males; white, dorsal, caudal tubercles; distinct black and white caudal bands variably present. These differences are summarized across all Southeast Asian species of Cnemaspis in Grismer et al. 2010 and 2014 (Tables 6,7).


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Comment 
EtymologyThe specific epithet honors Professor Shahrul Anuar Mohd Sah of Universiti Sains Malaysia for his extensive contributions to the understanding of the vertebrate zoology of Peninsular Malaysia as well as his active and scholarly mentorship of many students. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • GRISMER, L. LEE; P. L. WOOD, JR., EVAN S. H. QUAH, SHAHRUL ANUAR, EHWAN NGADI & NORHAYATI AHMAD 2015. A new insular species of Rock Gecko (Cnemaspis Boulenger) from Pulau Langkawi, Kedah, Peninsular Malaysia. Zootaxa 3985 (2): 203–218 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Grismer, L.L.; Chan Kin Onn; E. Quah, M.A. Muin; A.E. Savage; J.L. Grismer; N. Ahmad; LF. Greer III; A.C. Remegio 2010. Another new, diminutive Rock Gecko (Cnemaspis Strauch) from Peninsular Malaysia and a discussion of resource partitioning in sympatric species pairs. Zootaxa 2569: 55–66 - get paper here
  • Grismer, L.L.; PERRY JR L. WOOD, SHAHRUL ANUAR, AWAL RIYANTO, NORHAYATI AHMAD, MOHD A. MUIN, MONTRI SUMONTHA, JESSE L. GRISMER, CHAN KIN ONN, EVAN S. H. QUAH, OLIVIER S. A. PAUWELS 2014. Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia. Zootaxa 3880 (1): 001–147 - get paper here
  • Grismer, L.L.; PERRY L. WOOD JR., MAKETAB MOHAMED, KIN ONN CHAN, HEATHER M. HEINZ, ALEX S-I. SUMARLI, JACOB A. CHAN, ARIEL I. LOREDO 2013. A new species of karst-adapted Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) from a threatened karst region in Pahang, Peninsular Malaysia. Zootaxa 3746 (3): 463–472 - get paper here
  • Nguyen, H.N., Hung, C., Yang, M. et al. 2020. Sympatric competitors have driven the evolution of temporal activity patterns in Cnemaspis geckos in Southeast Asia. Sci Rep 10: 27 - get paper here
  • Onn, Chan Kin; L. L. Grismer, Shahrul Anuar, E. Quah, M.A. Muin, A.E. Savage, J.L. Grismer, Norhayati Ahmad, A.C. Remigio, L.F. Greer 2010. A new endemic rock Gecko Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Gunung Jerai, Kedah, northwestern Peninsular Malaysia. Zootaxa 2576: 59–68 - get paper here
  • QUAH, EVAN S.H.; SHAHRULANUAR, M.S., GRISMER, L.L., MOHD ABDUL MUIN, M.A.; CHAN, K.O. and GRISMER, J.L. 2011. Preliminary Checklist of the Herpetofauna of Jerejak Island, Penang, Malaysia. Malayan Nature Journal 63 (3): 595-600
  • Rooijen, J. van, Chan Kin Onn, L. Lee Grismer & Norhayati Ahmad 2011. Estimating the herpetofaunal species richness of Pangkor Island, Peninsular Malaysia. Bonn zoological Bulletin 60 (1): 3-8 - get paper here
  • WOOD, PERRY JR. L.; EVAN S.H. QUAH, SHAHRUL ANUAR M.S., MOHD ABDUL MUIN 2013. A new species of lowland karst dwelling Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from northwestern Peninsular Malaysia. Zootaxa 3691 (5): 538–558 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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