Cnemaspis shahruli GRISMER, CHAN, QUAH, MUIN, SAVAGE, GRISMER, AHMAD, GREER & REMEGIO, 2010
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Cnemaspis shahruli
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| Higher Taxa | Gekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) |
| Subspecies | |
| Common Names | E: Shahrul’s Rock Gecko |
| Synonym | Cnemaspis shahruli GRISMER, CHAN, QUAH, MUIN, SAVAGE, GRISMER, AHMAD, GREER & REMEGIO 2010 Cnemaspis shahruli — GRISMER 2011 Cnemaspis shahruli — GRISMER et al. 2014: 72 |
| Distribution | W Malaysia (Penang and Jerejak Islands, Penang; Sungai Sedim, Kedah; Pulau Pangkor, Perak) Type locality: Telok Bahang Recreational Forest Reserve, Penang (05°27.233’N 100°12.324’E), 67 m elevation. |
| Reproduction | oviparous (manual and phylogenetic imputation, fide Zimin et al. 2022) |
| Types | Holotype: ZRC 2.6898; adult male, collected on 14 March 2010 by Evan Quah. Paratypes. ZRC |
| Diagnosis | Diagnosis: Maximum SVL 36.5 mm; 10 or 11 supralabials; 8–10 infralabials; ventral scales keeled; no precloacal pores; 19–23 paravertebral tubercles; body tubercles randomly arranged, present on flanks; tubercles within lateral caudal furrows; ventrolateral caudal tubercles absent anteriorly; lateral row of caudal tubercles present anteriorly; caudal tubercles not encircling tail; all subcaudals keeled, no enlarged median row; 1–3 postcloacal tubercles on each side of tail base; no enlarged femoral or subtibial scales; subtibials keeled; usually no enlarged submetatarsal scales on first toe; 21–30 subdigital fourth toe lamellae; light-colored vertebral stripe variably present; gular region, throat and pectoral region yellow in males; dark, central, elongate marking occurs in mental region; a single ocellus in the shoulder region in males; white, dorsal, caudal tubercles; distinct black and white caudal bands variably present. These differences are summarized across all Southeast Asian species of Cnemaspis in Grismer et al. 2010 and 2014 (Tables 6,7). Comparisons. Cnemaspis shahruli sp. nov. can be separated from all species of Cnemaspis except C. siamensis (Smith) on the basis of having dark, longitudinal, mental markings. It can be separated even further from all other Cnemaspis except C. monachorum Grismer, Norhayati, Chan, Belabut, Muin, Wood & Grismer by having a maximum SVL of less than 39 mm. It differs from C. argus Dring, C. karsticola Grismer, Grismer, Wood & Chan, C. kumpoli Smith, C. mcguirei Grismer, Grismer, Wood & Chan, C. monachorum, C. nuicamensis Grismer & Ngo, C. pseudomcguirei, C. psychedelica Grismer, Ngo & Grismer, C. roticanai Grismer & Chan, C. siamensis, Cnemaspis vandeventeri Grismer, Sumontha, Cota, Grismer, Wood, Pauwels & Kunya; Cnemaspis kamolnorranathi Grismer, Sumontha, Cota, Grismer, Wood, Pauwels & Kunya; Cnemaspis huaseesom Grismer, Sumontha, Cota, Grismer, Wood, Pauwels & Kunya; and Cnemaspis punctatonuchalis Grismer, Sumontha, Cota, Grismer, Wood, Pauwels & Kunya in having 10 or more supralabials as opposed to nine or less. From C. affinis, C. aurantiacopes Grismer & Ngo, C. boulengeri Strauch, C. caudanivea Grismer & Ngo, C. chanthaburiensis Bauer & Das, C. flavigaster Chan & Grismer, C. tucdupensis Grismer & Ngo, Cnemaspis punctatonuchalis, and Cnemaspis niyomwanae, Grismer, Sumontha, Cota, Grismer, Wood, Pauwels & Kunya if differs in having keeled as opposed to smooth ventral scales. Cnemaspis shahruli lacks precloacal pores which differentiates it from C. bayuensis Grismer, Grismer, Wood & Chan, C. biocellata Grismer, Chan, Nasir & Sumontha, C. dringi Das & Bauer, C. flavolineata (Nicholls), C. nigridia (Smith), C. paripari Grismer & Chan, C. perhentianensis Grismer & Chan, Cnemaspis chanardi Grismer, Sumontha, Cota, Grismer, Wood, Pauwels & Kunya, and Cnemaspis narathiwatensis Grismer, Sumontha, Cota, Grismer, Wood, Pauwels & Kunya. It differs further from all species of Cnemaspis except C. affinis, C. biocellata, C. harimau Chan, Grismer, Shahrul, Quah, Muin, Savage & Grismer, C. kumpoli, C. mcguirei, and C. pseudomcguirei in having a black shoulder patch enclosing a single, white ocellus [from GRISMER et al. 2010]. |
| Comment | |
| Etymology | The specific epithet honors Professor Shahrul Anuar Mohd Sah of Universiti Sains Malaysia for his extensive contributions to the understanding of the vertebrate zoology of Peninsular Malaysia as well as his active and scholarly mentorship of many students. |
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