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Cophotis dumbara SAMARAWICKRAMA, RANAWANA, RAJAPAKSHA, ANANJEVA, ORLOV, RANASINGHE & SAMARAWICKRAMA, 2006

IUCN Red List - Cophotis dumbara - Critically Endangered, CR
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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Dumbara Agama
G: Dumbara-Agame 
SynonymCophotis dumbara SAMARAWICKRAMA, RANAWANA, RAJAPAKSHA, ANANJEVA, ORLOV, RANASINGHE & SAMARAWICKRAMA 2006
Cophotis dumbarae MANAMENDRA-ARACHCHI et al. 2006 
DistributionSri Lanka (Ceylon)

Type locality: Dothalugala Man and Biosphere Reserve, The Knuckles Forest Range, Central Province, Sri Lanka, 07°20’ 31.4” N, 80°50’ 25.6” E, in cardamon cultivated area, elevation 1425 m elevation  
Reproductionovoviviparous 
TypesHolotype: NMSL (Colombo National Museum) (2006 
DiagnosisDiagnosis: A new species of Cophotis can be distinguished from C. ceylanica by smaller scales of gular regions. Medium sized, slender Cophotis (SVL + TL up to
133.57 and 119.67 mm in males and females, respectively). Agamids with prehensile tail and not prominent tympanum. Number of scales around the midbody is 30. Lower surface of the head has smaller scales; their number (GS1) along the longitudinal axis of the gular sac is 34 – 35 and their number across the most wide part (GS2) of gular region is 23 – 24. Size of infralabials is smaller as well as size of all gular scales [from Samarawickrama et al. 2006]. 
CommentD. dumbara and C. dumbarae have been described independently. However, Hallermann & Böhme (2007) showed that the two species are identical and that C.dumbara has priority over C. dumbarae.

Habitat: arboreal; the species is mostly found on tree trunks. 
EtymologyNamed after the type locality. The Knuckles massif from which this species is recorded is separated from the rest of the central mountain region of Sri Lanka by a broad Dumbara Valley. 
References
  • Barts, M. & Kwet, A. 2007. Wettlauf um eine Baumagame - Cophotis dumbara oder Cophotis dumbarae? Draco 7 (30): 91-92 - get paper here
  • Hallermann, J. & W. Böhme 2007. Zur Validität von zwei kürzlich beschriebenen Arten der in Sri Lanka endemischen Gattung Cophotis PETERS, 1861 (Squamata: Agamidae) aus der Knuckles Forest Range. Salamandra 43 (3): 187-190 - get paper here
  • Manamendra-Arachchi, Kelum; Anslem de Silva and Thasun Amarasinghe 2006. Description of a second species of Cophotis (Reptilia: Agamidae) from the highlands of Sri Lanka. Lyriocephalus 6 (Suppl. 1): 1-8
  • Samarawickrama, V. A. M. P. K.; K. B. Ranawana, D. R. N. S. Rajapaksha, Natalia B. Ananjeva, Nikolai L. Orlov, J. M. A. S. Ranasinghe, and V. A. P.Samarawickrama 2006. A new species of the genus Cophotis (Squamata: Agamidae) from Sri Lanka. Russ. J. Herpetol. 13 (3): 207-214
  • Somaweera, R. & Somaweera, N. 2009. Lizards of Sri Lanka: a colour guide with field keys. Chimaira, Frankfurt, 304 pp.
  • SOYSA, H.D.C.; U.S.B. HETTIGE, M.G. COORAY & D.R. VIDANAPATHIRANE. 2021. Copulatory behavior of Knuckles pygmy lizard (Cophotis dumbara). Taprobanica 10(2): 128–130. - get paper here
 
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