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Crotalus intermedius TROSCHEL, 1865

IUCN Red List - Crotalus intermedius - Least Concern, LC
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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
SubspeciesCrotalus intermedius intermedius TROSCHEL in MÜLLER 1865
Crotalus intermedius gloydi TAYLOR 1941
Crotalus intermedius omiltemanus GÜNTHER 1895 
Common NamesE: Mexican Smallhead Rattlesnake
S: Viborilla 
SynonymCrotalus intermedius intermedius TROSCHEL in MÜLLER 1865
Crotalus intermedius TROSCHEL in MÜLLER 1865
Crotalus intermedius intermedius KLAUBER 1952: 9
Crotalus intermedius intermedius — LINER 1994
Crotalus intermedius — MCDIARMID, CAMPBELL & TOURÉ 1999: 286
Aechmophrys (Cottonus) intermedius — HOSER 2009
Crotalus intermedius — WALLACH et al. 2014: 191

Crotalus intermedius gloydi TAYLOR 1941
Crotalus triseriatus gloydi TAYLOR 1941
Crotalus gloydi lautus SMITH 1946
Crotalus intermedius gloydi DAVIS & DIXON 1957
Crotalus intermedius gloydi — LINER 1994
Crotalus intermedius gloydi — BRYSON 2007

Crotalus intermedius omiltemanus GÜNTHER 1895
Crotalus omiltemanus GÜNTHER 1895
Crotalus intermedius omiltemanus KLAUBER 1952: 14
Crotalus triseriatus BOULENGER 1896
Crotalus intermedius omiltemanus — LINER 1994
Crotalus intermedius omiltemanus — BRYSON 2007 
DistributionMexico (disjunct populations in SE Hidalgo, Puebla, SW Tlaxcala, WC Veracruz, Oaxaca, Guerrero)

intermedius: Mexico (Michoacan, NE Puebla, WC Veracruz, C Oaxaca)

gloydi: Mexico (C Oaxaca, Michoacan); Type locality: Mexico: Cerro San Felip, Oaxaca; E. H. Taylor; 1940.

omiltemanus: Mexico (C Guerrero)  
Reproductionovovivparous 
TypesHolotype: lost, originally in Müller collection, later in Bonn (ZFMK); destroyed in World War II, according to R. Mertens (pers. comm.) in Klauber (1952: 10).
Holotype: INHS (= UIMNH) 25070 (23645) [gloydi]
Syntypes: BMNH 1946.1.19.28-29 (both formerly BMNH 93.3.15.11). [omiltemanus]
Holotype: USNM 110598 [Crotalus gloydi lautus] 
DiagnosisAdditional details, e.g. a detailed description or comparisons (2646 characters), are available for collaborators and contributors. Please contact us for details. 
CommentVenomous!

Crotalus intermedius FISCHER 1882 is a primary homonym of Crotalus intermedius TROSCHEL in MÜLLER 1865 (Holotypus: UMB 435 missing and may be lost).

Nomenclature: Hoser’s 2009 classification and nomenclature has been rejected as unnecessary and unavailable by WÜSTER & BERNILS 2011. 
EtymologyNamed after Latin intermedius, that is between, intermediate. [“...Beschildung zu Crotalus s. str., wenngleich sie manche anderen Merkmale, namentlich die Theilung der letzten Subcaudalschilder mit Crotalophorus Gray gemein hat...”].

C. i. gloydi was named after Howard Kay Gloyd (1902-1978), US American herpetologist.

C. i. omiltemanus was named after Omilteme, a locality in Guerrero State, México + Latin -anus, pertaining to. ["...Hab. Mexico, Omilteme in Guerrero…"].
 
References
  • Alvarado-Diaz, J. & Campbell, J.A. 2004. A new montane rattlesnake (Viperidae) from Michoacán, Mexico. Herpetologica 60 (2): 281-286 - get paper here
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  • Bryson Jr, Robert W.; Robert W. Murphy,Matthew R. Graham, Amy Lathrop and David Lazcano 2011. Ephemeral Pleistocene woodlands connect the dots for highland rattlesnakes of the Crotalus intermedius group. Journal of Biogeography 38: - get paper here
  • Bryson, R.W. 2007. Fotopirsch auf Gebirgsklapperschlangen. Reptilia (Münster) 12 (66): 32-37 - get paper here
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