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Ctenophorus dualis (STORR, 1965)

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Spotted Sand-dragon, Spotted Dragon 
SynonymAmphibolurus maculatus dualis STORR 1965: 47
Phthanodon dualis — WELLS & WELLINGTON 1985: 18
Ctenophorus dualis — EDWARDS & HUTCHINSON 2023 
DistributionAustralia (SE Western Australia, extreme SW South Australia)

Type locality: 7 mi SSE of Cocklebiddy, in 32° 08’ S, 126° 08’ E, W. A.  
Reproductionoviparous 
TypesHolotype: WAM R23637 
DiagnosisDiagnosis. The largest of the C. maculatus complex. Gray males with orange to red dorsolateral stripes that are made more contrasting because of more or less continuous black dorsolateral and lateral coloring; females markedly larger than males with weakly contrasting whitish to grayish dorsolateral stripes. Male throat with black chevron markings and scattered additional dark speckling on the white gular region and black chest patch moderately well developed with a weak posteromedial extension (Figs. 5E–F, S8G, S9D–E). Femoral pore row reaches the knee. (Edwards & Hutchinson 2023) 
CommentDistribution: see map in EDWARDS & HUTCHINSON 2023: 177 (Fig. 1) 
EtymologyFrom the Latin dualis (of two kinds), in reference to the strong sexual dichromatism. (G. Shea, pers. comm., 9 Feb 2024) 
References
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Brygoo, E.R. 1988. Les types d’Agamidés (Reptiles, Sauriens) du Muséum national d’Histoire naturelle Catalogue critique. Bull. Mus. Nat. Hist. Nat. 10 (ser. 4) A (3), suppl.: 1-56
  • Chapple, David G.; Reid Tingley, Nicola J. Mitchell, Stewart L. Macdonald, J. Scott Keogh, Glenn M. Shea, Philip Bowles, Neil A. Cox, John C. Z. Woinarski 2019. The Action Plan for Australian Lizards and Snakes 2017. CSIRO, 663 pp. DOI: 10.1071/9781486309474 - get paper here
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Doughty, P.; Maryan, B.; Melville, J. & Austin, J. 2007. A new species of Ctenophorus (Lacertlia: Agamidae) from Lake Disappointment, Western Australia. Herpetologica 63 (1): 72-86 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Edwards, D., & Hutchinson, M. 2023. Sand Dragons: Species of the Ctenophorus maculatus Complex (Squamata: Agamidae) of Australia's Southern and Western Interior. Journal of Herpetology 57 (2): 176-196 - get paper here
  • Ellis, Ryan J. 2019. An annotated type catalogue of the dragon lizards (Reptilia: Squamata: Agamidae) in the collection of the Western Australian Museum. Records of the Western Australian Museum 34: 115–132 - get paper here
  • Escoriza Boj, D. 2005. Australia. Reptiles and Amphibians, Part 2: Desert and tropical savanna. Reptilia (GB) (41): 52-57 - get paper here
  • Glauert, L. 1959. Herpetological Miscellanea. XI. Dragon Lizards of the Genus Amphibolurus. Western Australian Naturalist 7 (2): 42-51 - get paper here
  • Gray, J. E. 1831. A synopsis of the species of Class Reptilia. In: Griffith, E & E. Pidgeon: The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with additional descriptions of all the species hither named, and of many before noticed [V Whittaker, Treacher and Co., London: 481 + 110 pp. [1830] - get paper here
  • Kinghorn, J. Roy 1924. Reptiles and batrachians from south and south-west Australia. Rec. Austral. Mus. 14 (3): 163-183 - get paper here
  • Licht, P., W.R. Dawson, V.H. Shoemaker & A.R. Main 1966. Observations on the thermal relations of Western Australian lizards Copeia 1966 (1): 97-110. - get paper here
  • Melville, Jane; Schulte, James A.; Larson, Allan 2001. A molecular phylogenetic study of ecological diversification in the Australian lizard genus Ctenophorus. Journal of Experimental Zoology: Molecular and Developmental Evolution 291(4): 339-353 - get paper here
  • Sternfeld, R. 1925. Beiträge zur Herpetologie Inner-Australiens. Abhandlungen Herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft, 38: 221—251
  • Storr, G. M. 1965. The Amphibolurus maculatus species-group (Lacertilia: Agamidae) in Western Australia. J R. Soc. West. Aust. 48: 45-54
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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