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Ctenophorus maculatus (GRAY, 1831)

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards) 
SubspeciesCtenophorus maculatus badius (STORR 1965)
Ctenophorus maculatus dualis (STORR 1965)
Ctenophorus maculatus griseus (STORR 1965)
Ctenophorus maculatus maculatus (GRAY 1831) 
Common NamesE: Spotted Sand-dragon, Spotted Dragon 
SynonymUromastyx maculatus GRAY 1831
Grammatophora Gaimardii DUMÉRIL & BIBRON 1837: 470
Amphibolurus maculatus — BOULENGER 1885: 381
Amphibolurus maculatus — KINGHORN 1924
Amphibolurus maculatus — COGGER 1983
Phthanodon maculatus — WELLS & WELLINGTON 1985: 19
Ctenophorus maculatus — COGGER 2000: 317
Ctenophorus maculatus — WILSON & SWAN 2010

Ctenophorus maculatus badius (STORR 1965)
Amphibolurus maculatus badius STORR 1965: 46
Phthanodon badius — WELLS & WELLINGTON 1985: 18

Ctenophorus maculatus dualis (STORR 1965)
Amphibolurus maculatus dualis STORR 1965: 47
Phthanodon dualis — WELLS & WELLINGTON 1985: 18

Ctenophorus maculatus griseus (STORR 1965)
Amphibolurus maculatus griseus STORR 1965: 47
Phthanodon griseus — WELLS & WELLINGTON 1985: 18 
DistributionAustralia (South Australia, Western Australia)

maculatus: west coast of Western Australia south of Shark Bay
badius: coast and interior of Western Australia between Shark Bay and Exmouth Gulf
griseus: SW interior of Western Australia
dualis: SE Western Australia and extreme SW South Australia.

Type locality: unknown, but probably Shark Bay, W. A. (fide COGGER 1983)
Type locality: 15 mi SE of Cardabia, in 23° 15’ S, 113° 55’ E, W. A. [badius]
Type locality: 7 mi SSE of Cocklebiddy, in 32° 08’ S, 126° 08’ E, W. A. [dualis]
Type locality: 14 mi W of Boorabbin, in 31° 15’ S, 120'05’ E, W. A. [griseus]  
Reproductionoviparous 
TypesHolotype: MNHN-RA 1259 (considered as nomen nudum by Brygoo agamid type catalog)
Holotype: WAM R18624 [badius]
Holotype: WAM R23637 [dualis]
Holotype: WAM R21708 [griseus] 
Diagnosis 
CommentGray's description referred to Cuvier's ms name applied to a specimen, almost certainly MNHP 1259, on which the name Grammatophora gaimardii is also based (COGGER 1983).
Habitat: shrubs, hummock grass. 
Etymology 
References
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Brygoo, E.R. 1988. Les types d’Agamidés (Reptiles, Sauriens) du Muséum national d’Histoire naturelle Catalogue critique. Bull. Mus. Nat. Hist. Nat. 10 (ser. 4) A (3), suppl.: 1-56
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp.
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Doughty, P.; Maryan, B.; Melville, J. & Austin, J. 2007. A new species of Ctenophorus (Lacertlia: Agamidae) from Lake Disappointment, Western Australia. Herpetologica 63 (1): 72-86 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Ellis, Ryan J. 2019. An annotated type catalogue of the dragon lizards (Reptilia: Squamata: Agamidae) in the collection of the Western Australian Museum. Records of the Western Australian Museum 34: 115–132 - get paper here
  • Escoriza Boj, D. 2005. Australia. Reptiles and Amphibians, Part 2: Desert and tropical savanna. Reptilia (GB) (41): 52-57 - get paper here
  • Glauert, L. 1959. Herpetological Miscellanea. XI. Dragon Lizards of the Genus Amphibolurus. Western Australian Naturalist 7 (2): 42-51
  • Gray, J. E. 1831. A synopsis of the species of Class Reptilia. In: Griffith, E & E. Pidgeon: The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with additional descriptions of all the species hither named, and of many before noticed [V Whittaker, Treacher and Co., London: 481 + 110 pp. [1830]
  • Kinghorn, J. Roy 1924. Reptiles and batrachians from south and south-west Australia. Rec. Austral. Mus. 14 (3): 163-183 - get paper here
  • Melville, Jane; Schulte, James A.; Larson, Allan 2001. A molecular phylogenetic study of ecological diversification in the Australian lizard genus Ctenophorus. Journal of Experimental Zoology: Molecular and Developmental Evolution 291(4): 339-353
  • Sternfeld, R. 1925. Beiträge zur Herpetologie Inner-Australiens. Abhandlungen Herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft, 38: 221—251
  • Storr, G. M. 1965. The Amphibolurus maculatus species-group (Lacertilia: Agamidae) in Western Australia. J R. Soc. West. Aust. 48: 45-54
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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