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Cubophis cantherigerus (BIBRON, 1840)

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Higher TaxaDipsadidae, Alsophiini, Colubroidea, Serpentes (snakes) 
SubspeciesCubophis cantherigerus adspersus (GUNDLACH & PETERS 1864)
Cubophis cantherigerus brooksi (BARBOUR 1914)
Cubophis cantherigerus cantherigerus (BIBRON 1843)
Cubophis cantherigerus pepei (SCHWARTZ & THOMAS 1970)
Cubophis cantherigerus schwartzi (LANDO & WILLIAMS 1969) 
Common NamesE: Cuban Racer
G: Kuba-Schlanknatter 
SynonymColuber cantherigerus BIBRON in DE LA SAGRA 1840: pl. 27
Dromicus angulifer BIBRON 1843: 133 (nom. subst.)
Dromicus angulifer — DUMÉRIL, BIBRON & DUMÉRIL 1854: 670
Alsophis angulifer var. caymanus — GARMAN 1887: 282
Dromicus angulifer — BOULENGER 1894: 120
Alsophis angulifer — KOOPMAN & RUIBAL 1955
Alsophis angulifer angulifer — SCHWARTZ & OGREN 1956
Alsophis cantherigerus spielmani GRANT 1959
Alsophis cantherigerus — SCHWARTZ & HENDERSON 1991: 569
Alsophis cantherigerus — CROTHER 1999
Alsophis cantherigerus — MATTISON 2007: 53
Ocyophis cantherigerus — ZAHER et al. 2009

Cubophis cantherigerus adspersus (GUNDLACH & PETERS 1864)
Dromicus (Alsophis) angulifer var. adspersus GUNDLACH & PETERS 1864: 388
Alsophis cantherigerus adspersus — BAUER et al. 1995: 71
Cubophis cantherigerus adspersus — HEDGES et al. 2009

Cubophis cantherigerus brooksi (BARBOUR 1914)
Alsophis cantherigerus brooksi — SCHWARTZ & HENDERSON 1991: 570
Cubophis cantherigerus brooksi — HEDGES et al. 2009

Cubophis cantherigerus cantherigerus (BIBRON 1843)
Alsophis cantherigerus cantherigerus — SCHWARTZ & HENDERSON 1991: 570
Cubophis cantherigerus cantherigerus — HEDGES et al. 2009

Cubophis cantherigerus pepei (SCHWARTZ & THOMAS 1970)
Alsophis cantherigerus pepei — SCHWARTZ & HENDERSON 1991: 571
Cubophis cantherigerus pepei — HEDGES et al. 2009

Cubophis cantherigerus schwartzi (LANDO & WILLIAMS 1969)
Alsophis cantherigerus schwartzi — SCHWARTZ & HENDERSON 1991: 571
Cubophis cantherigerus schwartzi — HEDGES et al. 2009 
DistributionCuba, Cayman Islands, Honduras (WILSON & MCCRANIE 2002)

brooksi: Swan Is.: Little Swan I.

pepei: Cuba: northern mesic coast of Guantanamo and Holguin provinces.

Type locality: “Cuba (Caimanera)” [adspersus] Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesSyntypes: MNHN 3545–46, MNHN 3561–63, and MCZ 2195 (formerly MNHN)
Syntypes: ZMB 5064A-B [adspersus]
HOLOTYPE: UIMNH 42341 [spielmani] 
CommentA number of subspecies of Alsophis cantherigerus have been elevated to full species status by HEDGES et al. 2009.

For illustrations see Grant, 1941; Peters and Schubert, 1968; Petzold, 1969; Maglio, 1970; Petzold, 1978; Kardong, 1980; Henderson and Sajdak, 1986.

Type species: Coluber cantherigerus BIBRON 1840 is the type species of the genus Cubophis HEDGES & VIDAL 2009.

Diagnosis (genus): Species in this genus have 17 midbody scale rows, 159–187 ventrals, 101–133 subcaudals, 2 apical scale pits, 8 upper labials, 10 lower labials, 13–17 total maxillary teeth, and 17–21 dentary teeth (Table 2). Cubophis differs in at least one of these characters from all other alsophiine genera except Hypsirhynchus. It differs from Hypsirhynchus in hemipenial characters such as the presence of enlarged papillate body calyces in the basal region and medial surface of the lobes (Zaher 1999). 
EtymologyEtymology (genus): The generic name refers to its distribution, centered on Cuba and nearby islands. 
References
  • Bauer,A.M.; Günther,R. & Klipfel,M. 1995. The herpetological contributions of Wilhelm C.H. Peters (1815-1883). SSAR Facsimile Reprints in Herpetology, 714 pp.
  • Boulenger, George A. 1894. Catalogue of the Snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Cocteau, J.-T. and G. Bibron 1838. Reptiles, in R. de la Sagra. Historia Física, Política y Natural de la Isla de Cuba, Tomo IV. Arthus Bertrand, Paris, 142 pp. [1838-1843 - index on page 143] - get paper here
  • Crother, Brian I. 1999. PHYLOGENETIC RELATIONSHIPS AMONG WEST INDIAN XENODONTINE SNAKES (SERPENTES; COLUBRIDAE) WITH COMMENTS ON THE PHYLOGENY OF SOME MAINLAND XENODONTINES. Contemporary Herpetology 2 - get paper here
  • Dominguez, Michel; Moreno, Luis V. 2006. Alsophis cantherigerus (Jubo, Jubo de Sabana, Jubo Sabanero). Size record. Herpetological Review 37 (3): 349. - get paper here
  • Estrada, A.R. 1998. Anfibios y Reptiles Encontrados Durante 1988 y 1989 en Cayo Paredón Grande, Archipiélago de Sabana-Camagüey, Cuba. Carib. J. Sci. 34 (1-2): 106-112
  • Fong G., Ansel and Gabriel Garcés G. 2002. Alsophis cantherigerus. Reptilia (GB) (21):55-57 - get paper here
  • Garman, S. 1887. On West Indian reptiles in the Museum of Comparative Zoology at Cambridge, Mass. Proc. Amer. Philos. Soc. 24: 278-286. - get paper here
  • Garman, S. 1887. On the reptiles and batrachians of Grand Cayman. Proc. Amer. Philos. Soc. 24: 273-277
  • Geißler, L. 2005. Exkursionen im Westen Kubas. Reptilia (Münster) 10 (56): 42-49 - get paper here
  • Grant, C. 1941. The herpetology of the Cayman Islands. Bulletin of the Institute of Jamaica, Science Series [1940] (2):1-56
  • Grant, C. 1959. New subspecies of Alsophis from Oriente, Cuba. Herpetologica 15 (1): 58-60 - get paper here
  • Hedges, S. Blair; Arnaud Couloux; Vidal, Nicolas 2009. Molecular phylogeny, classification, and biogeography of West Indian racer snakes of the Tribe Alsophiini (Squamata, Dipsadidae, Xenodontinae). Zootaxa 2067: 1-28 - get paper here
  • Henderson, R.W., and Sajdak, R.A. 1986. West Indian racers: disappearing act or a second chance?. Lore (Milwaukee Public Museum) 36(3):13-18.
  • Kardong, K.V. 1980. Jaw musculature of the West Indian snake Alsophis cantherigerus brooksi (Colubridae, Reptilia). Breviora (463):1-26. - get paper here
  • KLIMENT, P. 1993. Ein Beitrag zur Biologie der Kubanischen Natter Alsophis cantherigerus (BIBRON, 1840) Sauria 15 (4): 27-31 - get paper here
  • Koopman, K.F. & Ruibal, R. 1955. Cave-fossil vertebrates from Camaguey, Cuba. Breviora (46): 1-8 - get paper here
  • Maglio, Vincent J. 1970. West Indian Xenodontine colubrid snakes: Their probable origin, phylogeny, and zoogeography. Bull. Mus. Comp. Zool. Harvard 141 (1): 1-54 - get paper here
  • Mattison, Chris 2007. The New Encyclopedia of Snakes. Princeton University Press
  • McCranie J R 2011. The snakes of Honduras. SSAR, Salt Lake City, 725 pp.
  • Peters, G.,and Schubert, R. 1968. Theoretische und praktische Aspekte einer biologischen Expedition durch Kuba. Biol. Schule 8/9:337-46.
  • Peters,W. 1864. Über einige neue Säugethiere (Mormops, Macrotus, Vesperus, Molossus, Capromys), Amphibien (Plathydactylus, Otocryptis, Euprepes, Ungalia, Dromicus, Tropidonotus, Xenodon, Hylodes), und Fische (Sillago, Sebastes, Channa, Myctophum, Carassius, Barbus, Mber. k. preuss. Akad. Wiss. Berlin [1864]: 381-399 - get paper here
  • Petzold, H.G. 1969. Zur Haltung and Fortpflanzungsbiologieeiniger kubanischer Schlangen im Tierpark Berlin. Salamandra 5:124-40.
  • Petzold, H.G. 1978. Zur Haltung und Fortpflanzungsbiologie der Kuba-Schlanknatter, Alsophis cantherigerus (Bibron 1840), im Terrarium. Zool. Garten N.F., Jena 48(2/3):155-63.
  • Reyes Vázquez, Angel Eduardo, Yasit Segovia Vega, Ansel Fong G. 2013. Predation on birds by the Cuban racer Cubophis cantherigerus (Squamata: Dipsadidae) in Cuba. Herpetology Notes 6: 165-166 - get paper here
  • Rodríguez Schettino, Lourdes, Carlos A. Mancina & Vilma Rivalta González 2013. REPTILES OF CUBA: CHECKLIST AND GEOGRAPHIC DISTRIBUTIONS. Smithsonian Herp. Inf. Serv. (144): 1-96
  • Schmidt, D. & Kunz, K. 2005. Ernährung von Schlangen. Natur und Tier Verlag, Münster, 159 pp. - get paper here
  • Schwartz, A. and Ogren, L.H. 1956. A collection of reptiles and amphibians from Cuba, with the descriptionof two new forms. Herpetologica 12 (2): 91-110 - get paper here
  • Schwartz,A. & Henderson,R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Wilson, Larry David and James R. McCranie 2002. Update on the list of reptiles know from Honduras. Herpetological Review 33 (2):90-94 - get paper here
 
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