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Cyclemys enigmatica FRITZ, GUICKING, AUER, SOMMER, WINK & HUNDSDÖRFER, 2008

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Higher TaxaGeoemydidae (Geoemydinae), Testudinoidea, Testudines (turtles) 
Subspecies 
Common Names 
SynonymCyclemys enigmatica FRITZ, GUICKING, AUER, SOMMER, WINK & HUNDSDÖRFER 2008
Cyclemys enigmatica — CHAN-ARD et al. 2015 
DistributionIndonesia (Sumatra), Southern Malay Peninsula and Greater Sunda Islands

Type locality: Padang, Sumatra  
Reproductionoviparous. 
TypesHolotype: NMW 9811 (Naturhistorisches Museum Vienna = NHMW), coll. Consul J. Schild 1901, stuffed adult female). 
DiagnosisDiagnosis. Adult: Dark coloured, with mainly or entirely dark brown to black shell. Carapace ovoid. Femoral mid-seam longer than or equal to anal mid-seam; anal notch wide and obtuse- angled. Young adults with dense black radiating pattern on carapace and plastron; aged adults often with uniform dark brown carapace and black plastron. Head and neck stripes absent. Crown of the head uniform brown, in life distinctly lighter coloured than temporal region. Palate and tongue in alcohol pure white. Hatchling (Fig. 9): Underside of the shell brownish- yellow, mottled with small darker brown specks; head and neck uniform dark, without stripes. For maximum size and com- parison with diagnostic characters of other species, see Table 3 in FRITZ et al. 2008: 10. 
CommentSimilar species: has been confused with Heosemys grandis (e.g. in CHAN-ARD et al. 2015; the species in their Fig 2. is not C. enigmatica but Heosemys grandis, U. Fritz, pers. comm., 17. June 2012; Hikida 2015).

Habitat: freshwater (Rivers, swamps) 
EtymologyThe species name enigmatica, Latinized adjective from the Greek noun aínigma (enigma, conundrum or riddle) refers to the long-lasting confusion caused by the distinct morphology of the new species and its introgressed mitochondrial genome. 
References
  • Baker, Nick 2016. Dark-bellied Leaf Terrapin Cyclemys enigmatica at Pulau Langkawi, Peninsular Malaysia. SEAVR 2016: 1 - get paper here
  • Bour, R. 2008. Global diversity of turtles (Chelonii; Reptilia) in freshwater. Hydrobiologia 595:593–598
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Chan-ard, Tanya; Siriporn Thong-aree, Michael Cota and Sunchai Makchai 2011. Cyclemys of the Peat Swamp of Southernmost Thailand. The Thailand Natural History Museum Journal 5(2): 107-112
  • Fritz, U.; D Guicking, M Auer, R. S Sommer, M Wink, A. K Hundsdörfer 2008. Diversity of the Southeast Asian leaf turtle genus Cyclemys: how many leaves on its tree of life?. Zoologica Scripta doi:10.1111/j.1463-6409.2008.00332. - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Hikida, Tsutomu 2015. Book Review: A Field Guide to the Reptiles of Thailand. Nat. Hist. Bull. Siam Soc. 61(1): 41–51
 
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