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Higher TaxaGeoemydidae (Geoemydinae), Testudinoidea, Testudines (turtles) 
Common Names 
SynonymCyclemys enigmatica FRITZ, GUICKING, AUER, SOMMER, WINK & HUNDSDÖRFER 2008
Cyclemys enigmatica — CHAN-ARD et al. 2015 
DistributionIndonesia (Sumatra), Southern Malay Peninsula and Greater Sunda Islands

Type locality: Padang, Sumatra  
TypesHolotype: NMW 9811 (Naturhistorisches Museum Vienna = NHMW), coll. Consul J. Schild 1901, stuffed adult female). 
DiagnosisDiagnosis in FRITZ et al. 2009.
CommentSimilar species: has been confused with Heosemys grandis (e.g. in CHAN-ARD et al. 2015; the species in their Fig 2. is not C. enigmatica but Heosemys grandis, U. Fritz, pers. comm., 17. June 2012; Hikida 2015).

Habitat: freshwater (Rivers, swamps) 
EtymologyThe species name enigmatica, Latinized adjective from the Greek noun aínigma (enigma, conundrum or riddle) refers to the long-lasting confusion caused by the distinct morphology of the new species and its introgressed mitochondrial genome. 
  • Baker, Nick 2016. Dark-bellied Leaf Terrapin Cyclemys enigmatica at Pulau Langkawi, Peninsular Malaysia. SEAVR 2016: 1 - get paper here
  • Bour, R. 2008. Global diversity of turtles (Chelonii; Reptilia) in freshwater. Hydrobiologia 595:593–598
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Chan-ard, Tanya; Siriporn Thong-aree, Michael Cota and Sunchai Makchai 2011. Cyclemys of the Peat Swamp of Southernmost Thailand. The Thailand Natural History Museum Journal 5(2): 107-112
  • Fritz, U.; D Guicking, M Auer, R. S Sommer, M Wink, A. K Hundsdörfer 2008. Diversity of the Southeast Asian leaf turtle genus Cyclemys: how many leaves on its tree of life?. Zoologica Scripta doi:10.1111/j.1463-6409.2008.00332. - get paper here
  • Hikida, Tsutomu 2015. Book Review: A Field Guide to the Reptiles of Thailand. Nat. Hist. Bull. Siam Soc. 61(1): 41–51
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