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Higher TaxaGeoemydidae (Geoemydinae), Testudinoidea, Testudines (turtles)
Common Names 
Cyclemys gameli – DAS et al. 2009 (in error)
Cyclemys gemeli — TTWG 2014: 377
Cyclemys gemeli — TTWG 2021
Cyclemys gemelli — MALSAWMDAWNGLIANA et al. 2022 (in error) 
DistributionNE India (Assam, Arunachal Pradesh, Nagaland), Bhutan, Nepal

Type locality: street from Tezpur to Arunachal Pradesh, 5 km to border of Arunachal Pradesh, Jia Bhoroli River Region, Assam, India.  
TypesHolotype: NMW 37153, Naturhistorisches Museum Vienna (NHMW), coll. R. Gemel 7 November 2003, incomplete bony shell of adult, sex unknown). 
DiagnosisDiagnosis. Adult: Dark coloured, resembling the other dark-bellied Cyclemys species but carapace elongated to rectangular and crown of the head uniform brown, not lighter coloured than temporal region. Hatchling: Unknown. For maximum size and comparison with diagnostic characters of other species, see Table 3 in Fritz et al. 2008. 
CommentHabitat: freshwater (Rivers, swamps) 
EtymologyThe species is named in honour of the collector of the holotype, Richard Gemel (Vienna), in recognition of his work in the Herpetological Collection of the Natural History Museum Vienna. 
  • Basumatary, Rajeev and D.K. Sharma 2013. The turtle fauna of Kaziranga National Park, Assam, India with notes on natural history and conservation status. Herpetology Notes 6: 59-72
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Bhupathy, S., Ramesh Kumar, S., Paramanandham, J., Thirumalainathan, P. and Pranjit Kumar Sarma 2013. Conservation of reptiles in Nagaland, India. In: K. K. Singh et al. (editors), Bioresources and Traditional Knowledge of Northeast India, Mizo Post-Graduate Science Society, pp. 181-186
  • Bour, R. 2008. Global diversity of turtles (Chelonii; Reptilia) in freshwater. Hydrobiologia 595:593–598 - get paper here
  • Das, Abhijit; Uttam Saikia, B. H. C. K. Murthy, Santanu Dey and Sushil K. Dutta 2009. A herpetofaunal inventory of Barail Wildlife Sanctuary and adjacent regions, Assam, north-eastern India. Hamadryad 34 (1): 117 – 134 - get paper here
  • Das, Indraneil & Abhijit Das 2017. A Naturalist’s Guide to the Reptiles of India, Bangladesh, Bhutan, Nepal, Pakistan and Sri Lanka. John Beaufoy Publishing Ltd., Oxford, 176 pp.
  • Fritz, U.; D Guicking, M Auer, R. S Sommer, M Wink, A. K Hundsdörfer 2008. Diversity of the Southeast Asian leaf turtle genus Cyclemys: how many leaves on its tree of life? Zoologica Scripta doi:10.1111/j.1463-6409.2008.00332. - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Kundu, Shantanu; Vikas Kumar, Boni Amin Laskar, Kailash Chandra, Kaomud Tyagi 2016. Mitochondrial DNA effectively detects non-native Testudines: Invisible wildlife trade in northeast India. Gene Reports, doi:10.1016/j.genrep.2016.02.002 - get paper here
  • Malsawmdawngliana, B. Boruah, N.G. Patel, S. Lalronunga, I. Zosangliana, K. Lalhmangaiha & A. Das 2022. An updated checklist of reptiles from Dampa Tiger Reserve, Mizoram, India, with sixteen new distribution records. Journal of Threatened Taxa 14(10): 21946–21960 - get paper here
  • PRASCHAG, PETER; ANNA K. HUNDSDÖRFER & UWE FRITZ 2009. Further specimens and phylogenetic position of the recently described leaf turtle species Cyclemys gemeli (Testudines: Geoemydidae). Zootaxa 2008: 29-37 - get paper here
  • Purkayastha J. 2013. An Amateur’s Guide to Reptiles of Assam. EBH Publishers (India) - get paper here
  • Purkayastha, J., DAS, I., Sengupta, S. 2015. Freshwater Turtles & Tortoises of South Asia. Bhabani Books, 138 pp. - get paper here
  • Purkayastha, Jayaditya; Ahmed Mahmadul Hassan, Hasanul Islam, Jessica Das, Manoj Sarma, Mituseela Basumatary, Nilakshi Sarma, Nishant Chatterjee, Sachin Singha, Vishnupriya Nair, Arundhati Purkayastha, Jayashree Dutta, Madhurima Das 2013. Turtles of the Temple Pond of Kamakhya, Assam, India. Reptile Rap (15): 11-15 - get paper here
  • Sinha, B., & Nath, K. P. 2022. First confirmed report of Assam Leaf Turtle, Cyclemys gemeli Fritz et al., 2008 (Testudines: Testudinoida: Geoemydidae) from Arunachal Pradesh, India. Records of the Zoological Survey of India, 122(1): 97-100 - get paper here
  • THAPA, AVANTIKA; PUJAN PRADHAN, MUKESH THAKUR, BHEEM DUTT JOSHI, LALIT KUMAR SHARMA & KAILASH CHANDRA. 2021. A new elevational and locality record of the Assam Leaf Turtle, Cyclemys gemeli (Geoemydidae), from the Darjeeling Hills, in the Central Himalayan Biogeographic Province. Reptiles & Amphibians 28(2): 326-328 - get paper here
  • Thapa, Avantika; Pujan Pradhan, Mukesh Thakur, Bheem Dutt Joshi, Lalit Kumar Sharma, Kailash Chandra 2021. A new elevational and locality record of the Assam Leaf Turtle, Cyclemys gemeli (Geoemydidae), from the Darjeeling Hills, in the Central Himalayan Biogeographic Province. Reptiles & Amphibians 28 (2): 326-328 - get paper here
  • TTWG [Peter Paul van Dijk, John B. Iverson, Anders G.J. Rhodin, H. Bradley Shaffer, and Roger Bour] 2014. Turtles of the World, 7th Edition: Annotated Checklist of Taxonomy, Synonymy, Distribution with Maps, and Conservation Status. 000.v7. Chelonian Research Monographs (ISSN 1088-7105) No. 5, doi:10.3854/crm.5.000.checklist.v7.2014 - get paper here
  • TTWG; Rhodin, A.G.J., Iverson, J.B., Bour, R., Fritz, U., Georges, A., Shaffer, H.B., and van Dijk, P.P. 2021. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (9th Ed.). In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Stanford, C.B., Goode, E.V., Buhlmann, K.A., and Mittermeier, R.A. (Eds.). Chelonian Research Monographs 8:1–472. doi:10.3854/crm.8.checklist.atlas.v9.2021. - get paper here
  • Wangyal, Jigme Tshelthrim; Dorji Wangchuk, and Indraneil Das 2012. First Report of Turtles from the Himalayan Kingdom of Bhutan. Chelonian Conservation and Biology Dec 2012, Vol. 11, No. 2: 268-272. - get paper here
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