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Cyrtodactylus celatus KATHRINER, BAUER, O’SHEA, SANCHEZ & KAISER, 2014

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus celatus KATHRINER, BAUER, O'SHEA, SANCHEZ & KAISER 2014
Gymnodactylus (?) marmoratus (KUHL) — SMITH 1927: 201
Gymnodactylus marmoratus DUMÉRIL & BIBRON (sic) — MERTENS 1930: 239 (part)
Gymnodactylus wetariensis Dunn — BRONGERSMA 1953: 174 (part)
Cyrtodactylus marmoratus — BAUER & HENLE 1994: 27 (part)
Cyrtodactylus celatus — RÖSLER 2016: 11 
DistributionIndonesia (West Timor)

Type locality: Djamplong, 55 kilometres by road from Kupang,” now Tjamplong (or Camplong), West Timor, Nusa Tenggara Timur Province, Indonesia  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: BMNH 1926.10.30.45 (Figs. 2–5), an adult female collected by Malcolm Smith between 26 February and 29 April 1924. 
DiagnosisDiagnosis: Cyrtodactylus celatus is a small (SVL of only known specimen 38.4 mm) species of Cyrtodactylus, distinguished from all other Sunda Shelf species by having large, elongate, conical, keeled tubercles on body, limbs, and tail; elongate, tubercles absent on the ventrolateral body fold and on the ventrolateral margin of the tail; 16 longitudinal rows of tubercles at midbody; 42 ventral scales between the ventrolateral folds; no transversely enlarged, median, subcaudal scales, uniform granular caudal scales; proximal subdigital lamellae transversely expanded; 17 subdigital lamellae (seven basal + ten distal) on the fourth toe; no abrupt transition between postfemoral and ventral femoral scales; no digital webbing; tail round in cross-section; ventrolateral caudal fringe lacking; no white reticulations on the head; and faint, squarish blotches on the body. 
CommentAbundance: only known from the type specimen (Meiri et al. 2017). 
EtymologyThe species name celatus is a Latin adjective, meaning ‘hidden away.’ We use it to describe this species not only because the specimen remained hidden behind a single line of Smith’s (1927) contribution and on the shelves of the BMNH, but also because it has taken nearly 200 years since the initial herpetological survey work on Timor to describe the first putatively endemic member of this diverse gecko genus from that island. 
References
  • Bauer, A.M. 1994. Liste der rezenten Amphibien und Reptilien: Gekkonidae I (Australia). Das Tierreich, Vol. 108, W. de Gruyter & Co. (Berlin)
  • Brongersma, L.D. 1953. Gymnodactylus marmoratus (Gray). Proceedings, Koninklijke Nederlandsche Akademie van Wetenschappen, 62: 172–175
  • HARTMANN, LUKAS; SVEN MECKE, MAX KIECKBUSCH, FELIX MADER, HINRICH KAISER 2016. A new species of bent-toed gecko, genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with taxonomic remarks on C. fumosus (Müller, 1895). Zootaxa 4067 (5): 552–568 - get paper here
  • KATHRINER, ANDREW; AARON M. BAUER, MARK O’SHEA, CAITLIN SANCHEZ, HINRICH KAISER 2014. Hiding in plain sight: a new species of bent-toed gecko (Squamata: Gekkonidae: Cyrtodactylus) from West Timor, collected by Malcolm Smith in 1924. Zootaxa 3900 (4): 555–568 - get paper here
  • Kemp, Christopher. 2017. The Lost Species: Great Expeditions in the Collections of Natural History Museums. University of Chicago Press, Chicago, xxii, 250 pp. [ISBN: 9780226386218]
  • MECKE, SVEN; MAX KIECKBUSCH, LUKAS HARTMANN, HINRICH KAISER 2016. Historical considerations and comments on the type series of Cyrtodactylus marmoratus Gray, 1831, with an updated comparative table for the bent-toed geckos of the Sunda Islands and Sulawesi. Zootaxa 4175 (4): 353–365 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Mertens, R. 1930. Die Amphibien und Reptilien der Inseln Bali, Lombok, Sumbawa und Flores. Senck. Naturf. Gesell., Frankfurt am Main, Abhandl. 42(3): 117-344.
  • O’SHEA, Mark; Caitlin SANCHEZ, Andrew KATHRINER, Sven MECKE, Venancio LOPES CARVALHO, Agivedo VARELA RIBEIRO, Zito AFRANIO SOARES, Luis LEMOS DE ARAUJO and Hinrich KAISER 2015. Herpetological Diversity of Timor-Leste: Updates and a Review of Species Distributions. Asian Herpetological Research 6 (2): 73–131 - get paper here
  • Rösler H 2016. Bemerkungen über einige Geckoarten der Zoologischen Staatssammlung München. Gekkota Suppl. 3: 1-94
  • Rösler, Herbert & Hinrich Kaiser 2016. Male secondary sexual characteristics of the gecko Cyrtodactylus celatus Kathriner et al., 2014 from Timor Island (Squamata, Gekkonidae). Spixiana 39 (1): 125-129 - get paper here
  • Smith, M. A. 1927. Contribution to the herpetology of the Indo-Australian Region. Proc. Zool. Soc. London, 1: 199-225.
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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