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Cyrtodactylus batucolus GRISMER, CHAN, GRISMER, WOOD & BELABUT, 2008

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Besar Island Bent-toed Gecko 
SynonymCyrtodactylus batucolus GRISMER, CHAN, GRISMER, WOOD & BELABUT 2008
Cyrtodactylus batucolus — GRISMER 2011
Cyrtodactylus batucolus — GRISMER & QUAH 2019 
DistributionW Malaysia (Pulau Besar, Melaka)

Type locality: Panti Putera (02°06.581 N, 102°19.757 E; 39 m elevation), Pulau Besar, Melaka, Peninsular Malaysia.  
Reproductionoviparous 
TypesHolotype: ZRC 2.6743, adult male, collected by Jesse L. Grismer, Chan K. Onn, Perry L. Wood, Jr., and L. Lee Grismer on 5 June 2008. 
DiagnosisDiagnosis: Cyrtodactylus batucolus is distinguished from all other Sunda Shelf species by having a maximum SVL of 75.2 mm; moderately sized, conical, keeled, body tubercles; tubercles occurring on occiput, forelimbs, hind limbs, and beyond base of tail; 38–42 ventral scales; no transversely enlarged, median, subcaudal scales; proximal subdigital lamellae transversely expanded; 17–19 subdigital lamellae on fourth toe; abrupt transition between postfemoral and ventral femoral scales; enlarged femoral and precloacal scales with a continuous series of 43–46 pore-bearing scales in males; precloacal groove absent, precloacal depression present; no white reticulum on head; dark blotches on body. These characters are summarized across all Sunda Shelf species in Table 1 in GRISMER et al. 2008. 
CommentAbundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet batucolus comes from the Malaysian word batu meaning rock and the Latin suffix -cola meaning to inhabit or to dwell in. 
References
  • Amarasinghe, A. A.; Awal Riyanto, Mumpuni & Lee L. Grismer 2020. A NEW BENT-TOED GECKO SPECIES OF THE GENUS Cyrtodactylus GRAY, 1827 (SQUAMATA: GEKKONIDAE) FROM THE WEST BALI NATIONAL PARK, BALI, INDONESIA. TAPROBANICA 9 (1): 59–70, pls. 22–24. - get paper here
  • Goldberg, S. R. and L. L. Grismer 2016. Cyrtodactylus batucolus (Besar Island Bent-toed Gecko) reproduction. Herpetological Review 47(3): 464. - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Grismer, L. Lee & Ahmad, N. 2008. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa 1924: 53–68 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L. Lee; Chan K. Onn; Grismer, J.L.; Wood, P.L. & Belabut, D. 2008. Three new species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia. Zootaxa 1921: 1–23 - get paper here
  • Grismer, L. Lee; Perry L. Wood, Jr. and Kelvin K. P. Lim 2012. Cyrtodactylus majulah, a new species of bent-toed gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology 60 (2): 487-499 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Onn, C.K.; Griserm, L.L.; Wood, P.L.; Grismer, J.L. & Ahmad, N. 2009. Preliminary Checklist of the Herpetofauna of Pulau Besar, Melaka, Malaysia. Tropical Life Sciences Research, 20(1): 81–87
  • Onn, Chan Kin & Norhayati Ahmad 2010. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa 2389: 47–56 - get paper here
 
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