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Cyrtodactylus payacola JOHNSON, QUAH ANUAR, MUIN, WOOD, GRISMER, GREER, ONN, AHMAD, BAUER & GRISMER, 2012

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus payacola JOHNSON, QUAH ANUAR, MUIN, WOOD, GRISMER, GREER, ONN, AHMAD, BAUER & GRISMER 2012
Cyrtodactylus payacola — GRISMER & QUAH 2019 
DistributionWest Malaysia (Penang)

Type locality: Bukit Panchor State Park, Penang, West Malaysia 05°09.465' N, 100°32.885' E at an elevation of 47 m.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: LSUHC 10074, adult male, collected by E. Quah S. H. on 4 September 2011. Paratypes. All paratopotypes were collected in various combinations by M. A. Muin, E. Quah S.H., S. Anuar, C. K. Onn, and L. L. Grismer from the same locality as the holotype. LSUHC 9982 was collected on 6 March 2010; LSUHC 10070 and 10071 were collected on 29 June 2011. 
DiagnosisDiagnosis: Cyrtodactylus payacola sp. nov. is distinguished from all other Sundaland species by having a maximum SVL of 67.7 mm; moderately sized, conical, keeled body tubercles; tubercles occurring on the occiput, forelimbs, hind limbs, and beyond base of tail; 44–51 ventral scales; no transversely enlarged, median subcaudal scales; 18–21 subdigital lamellae on fourth toe; abrupt transition between postfemoral and ventral femoral scales; no femoral pores; 11 or 12 contiguous, pore-bearing precloacal scales; shallow, longitudinal, precloacal groove; a pair of posteromedially elongate, triangular to semilunar-shaped, paravertebral blotches on nape prominently out- lined in light yellow; no wide, dark, ventrolateral stripes on flanks; no white reticulum on top of head; paired, semi- transversely arranged, dark blotches on body.


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Comment 
EtymologyThe specific epithet payacola is derived from the word paya meaning “swamp” in the Malay language and the Latin suffix colo, which means to inhabit or dwell in and is in reference to microhabitat in which this species is found. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L. Lee; Perry L. Wood, Jr. and Kelvin K. P. Lim 2012. Cyrtodactylus majulah, a new species of bent-toed gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology 60 (2): 487-499 - get paper here
  • Grismer, Lee; Wood, Perry L., Jr.; Poyarkov, Nikolay A.; Le, Minh D.; Karunarathna, Suranjan; Chomdej, Siriwadee; Suwannapoom, Chatmongkon; Qi, Shuo; Liu, Shuo; Che, Jing; Quah, Evan S.H.; Kraus, Fred; Oliver, Paul M.; Riyanto, Awal; Pauwels, Olivier 2021. Karstic Landscapes Are Foci of Species Diversity in the World’s Third-Largest Vertebrate Genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata; Gekkonidae). Diversity 13 (5): 183 - get paper here
  • HARVEY, MICHAEL B.; KYLE A. O’CONNELL, GABRIEL BARRAZA, AWAL RIYANTO, NIA KURNIAWAN & ERIC N. SMITH 2015. Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the Southern Bukit Barisan Range of Sumatra and an estimation of their phylogeny. Zootaxa 4020 (3): 495–516 - get paper here
  • JOHNSON, CHELSÉA B.; EVAN QUAH, S. H., SHAHRUL ANUAR, M. A. MUIN, PERRY L. WOOD, JR., JESSE L. GRISMER, LEE F. GREER, CHAN KIN ONN, NORHAYATI AHMAD, AARON M. BAUER & L. LEE GRISMER 2012. Phylogeography, geographic variation, and taxonomy of the Bent-toed Gecko Cyrtodactylus quadrivirgatus Taylor, 1962 from Peninsular Malaysia with the description of a new swamp dwelling species. Zootaxa 3406: 39–58 - get paper here
  • Riedel, J., Eisele, K., Gabelaia, M., Higham, T. E., Wu, J., Do, Q. H., Nguyen, T. Q., Meneses, C. G., Brown, R. M., Ziegler, T., Grismer, L. L., Russell, A. P., & Rödder, D. 2024. Ecologically-related variation of digit morphology in Cyrtodactylus (Gekkota, Squamata) reveals repeated origins of incipient adhesive toepads. Functional Ecology, 00, 1–19 - get paper here
  • RIYANTO, AWAL; L. LEE GRISMER & PERRY L. WOOD, JR. 2015. Cyrtodactylus rosichonariefi sp. nov. (Squamata: Gekkonidae), a new swamp-dwelling bent-toed gecko from Bunguran Island (Great Natuna), Indonesia. Zootaxa 3964 (1): 114–124 [erratum in Zootaxa 3999 (4): 600] - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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