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Cyrtodactylus psarops HARVEY, O'CONNELL, BARRAZA, RIYANTO, KURNIAWAN & SMITH, 2015

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Speckle-faced Bent-toed Geckos
Indonesian: Cicak Jari Lengkung Muka Bintik 
SynonymCyrtodactylus psarops HARVEY, O'CONNELL, BARRAZA, RIYANTO, KURNIAWAN & SMITH 2015
Cyrtodactylus cf. psarops — FAUZI & KURNIAWAN 2018 
DistributionIndonesia (S Sumatra)

Type locality: Gunung Tanggamus, above Gisting, 5.424° S, 104.692° N, 1163 m, Kabupaten Tanggamus, Lampung Province, Sumatra, Indonesia  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: MZB 9699, adult female; field number ENS 14612, Fig. 1), collected June 11, 2013, by M. B. Harvey, E. N. Smith, and field parties from BC, MZB, UB, and UTA.
Paratypes (14). Four males (MZB 9686–9687, MZB 9700, UTA R-61589) and one unsexed juvenile (UTA R- 61579) from forest above Ngarip, 5.280–5.284° S, 104.557–104.558°E, 1318–1419 m; four females (UTA R- 61580–61581, MZB 9690–9691) and one male (UTA R-61590) from forest NE of Ngarip, 5.298-5.314° S, 104.528–104.546, 950–1065 m; one male (MZB 9697) and two females (UTA R-61582–61583) from Gunung Tanggamus, above Gisting, 5.422–5.424° S, 104.692–104.695° N, 1016–1163 m, Lampung, Sumatra, Indonesia; and one female (MZB 9700) from Gunung Pesagi (known locally as “Masagi”), Remanan Jaya, Muara dua, Sumatera Selatan Province, Sumatra, Indonesia. M. B. Harvey, E. N. Smith, and field parties from BC, MZB, UB, and UTA collected the paratypes on 9–17 June 2013. 
DiagnosisDiagnosis: A medium-size species of Cyrtodactylus reaching 82 mm SVL and distinguished from all congeners by the following combination of characters: (1) body moderately robust; limbs and digits relatively long; (2) tail 1.06–1.35 times longer than body; (3) subpyramidal, weakly keeled tubercles extending from frontal region and supraorbital skin to base of tail, 23–26 irregular longitudinal rows of tubercles at midbody, 28–38 irregular transverse rows between limbs; (4) dorsal antebrachium and often brachium tuberculate; (5) tubercles on tail restricted to basal third; (6) 38–49 ventrals in a transverse row at midbody; (7) subcaudals not transversely enlarged; (8) deep, subtriangular depression in precloacal region of males; (9) 28–32 femoral and precloacal pores in males, arrayed in a continuous series (fewer pore primordia usually present in females), precloacal pores sunken into precloacal depression; (10) scale at anterior apex of pore-bearing scales greatly enlarged, sunken into precloacal depression in males, superficial in females; (11) pore-bearing series sharply transitioning to tiny granular scales of postaxial surface of thigh; (12) 18–22 lamellae under fourth toe; (13) cloacal tubercles usually single; (14) dorsal pattern of brown blotches edged first in dark brown to black then in pale gray or brown; at least some tubercles on flanks yellow to pale cream; (15) labials charcoal or dark brown with yellow or cream spots; occiput with few large blotches; long postocular stripe extending beyond arm onto flank; (16) 7–9 transverse, irregular or broken bands on body; 10–13 regular bands on tail, mostly complete ventrally.
 
Comment 
EtymologyThis species’ name psarops is a noun in apposition derived from the Greek words psaros, meaning speckled, and ops, meaning face. The new name alludes to the characteristic yellow speckling of the labials and snout in this species. 
References
  • Fauzi, M A; N Kurniawan 2018. Classification of Cyrtodactylus marmoratus species complex based on the molecular approach. Journal of Physics: Conf. Series 1146 (2019) 012040 - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • HARTMANN, LUKAS; SVEN MECKE, MAX KIECKBUSCH, FELIX MADER, HINRICH KAISER 2016. A new species of bent-toed gecko, genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with taxonomic remarks on C. fumosus (Müller, 1895). Zootaxa 4067 (5): 552–568 - get paper here
  • HARVEY, MICHAEL B.; KYLE A. O’CONNELL, GABRIEL BARRAZA, AWAL RIYANTO, NIA KURNIAWAN & ERIC N. SMITH 2015. Two new species of Cyrtodactylus (Squamata: Gekkonidae) from the Southern Bukit Barisan Range of Sumatra and an estimation of their phylogeny. Zootaxa 4020 (3): 495–516 - get paper here
  • MECKE, SVEN; MAX KIECKBUSCH, LUKAS HARTMANN, HINRICH KAISER 2016. Historical considerations and comments on the type series of Cyrtodactylus marmoratus Gray, 1831, with an updated comparative table for the bent-toed geckos of the Sunda Islands and Sulawesi. Zootaxa 4175 (4): 353–365 - get paper here
  • O’Connell, K. A., Smart, U., Sidik, I., Riyanto, A., Kurniawan, N., & Smith, E. N. 2019. Diversification of bent-toed geckos (Cyrtodactylus) on Sumatra and west Java. Molecular Phylogenetics and Evolution, 134, 1–11 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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