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Cyrtodactylus majulah GRISMER, WOOD & LIM, 2012

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common Names 
SynonymCyrtodactylus majulah GRISMER, WOOD & LIM 2012
Gymnodactylus marmoratus – SWORDER 1925: 63 (not of KUHL)
Gymnodactylus marmoratus – SMITH 1930: 12
Cyrtodactylus quadrivirgatus – LIM & LIM 1992: 121 (not of TAYLOR)
Cyrtodactylus quadrivirgatus – BAKER & LIM 2008: 80
Cyrtodactylus quadrivirgatus – DRING 1979: 228 (in part)
Cyrtodactylus quadrivirgatus – DENZER & MANTHEY 1991: 314
Cyrtodactylus quadrivirgatus – CHAN-ARD et al., 1999: 24
Cyrtodactylus quadrivirgatus – MANTHEY & GROSSMANN 1997: 227 
DistributionSingapore (Central Catchment Nature Reserve); Indonesia (Pulau Bintan in the Riau Archipelago)

Type locality: Singapore: Central Catchment Nature Reserve, Nee Soon Swamp-forest (1°22'48.99"N, 103°49'05.39"E; 51 m elevation).  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZRC 2.6950, adult male, 67.9 mm SVL; coll: L. Grismer, E. Quah, K. O. Chan, A. Figueroa & A. Devan-Song, 22 Jun.2011. Paratypes. — Adult male: 68.0 mm SVL (ZRC 2.6951); adult male: 62.7 mm SVL (ZRC 2.6952); adult male: 67.2 mm SVL (LSUHC 10458); adult male: 67.2 mm SVL (ZRC 2.6953); collection data as per holotype. 
DiagnosisDiagnosis: — Cyrtodactylus majulah new species is differentiated from its Sundaland congeners by having the unique combination of a maximum SVL of 68.0 mm; eight or nine supralabials; seven or eight infralabials; strongly tuberculated body and limbs; 39–46 paravertebral tubercles; 43–53 ventral scales; no abrupt contact of large and small postfemoral scales; no enlarged femoral scales; 20–23 subdigital lamellae; no femoral pores; no deep precloacal groove; enlarged precloacal scales; 7–11 precloacal pores; no enlarged median subcaudals; tubercles on the anterior portion of the tail; no reticulated pattern on head; having a blotched dorsal pattern; and lacking the paired, dark, semi-lunar shaped blotches on the upper nape prominently outlined in white. These and other characters are scored across all Sundaland species (Table 4). 
CommentSynonymy: after GRISMER et al. 2012. 
References
  • FIGUEROA, ALEX; MARTYN E.Y. LOW, KELVIN K.P. LIM 2023. Singapore’s herpetofauna: updated and annotated checklist, history, conservation, and distribution. Zootaxa 5287 (1): 001–378 - get paper here
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • Grismer, L. Lee; Perry L. Wood, Jr. and Kelvin K. P. Lim 2012. Cyrtodactylus majulah, a new species of bent-toed gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology 60 (2): 487-499 - get paper here
  • Leo S, Suherman M, Permatasari A, Suganda D, Zulamri, Winarni NL. 2020. Herpetofauna diversity in Zamrud National Park, Indonesia: baseline checklist for a Sumatra peat swamp forest ecosystem. Amphibian & Reptile Conservation 14(2) [General Section]: 250–263 (e249) - get paper here
  • RIYANTO, AWAL; L. LEE GRISMER & PERRY L. WOOD, JR. 2015. Cyrtodactylus rosichonariefi sp. nov. (Squamata: Gekkonidae), a new swamp-dwelling bent-toed gecko from Bunguran Island (Great Natuna), Indonesia. Zootaxa 3964 (1): 114–124 [erratum in Zootaxa 3999 (4): 600] - get paper here
  • Teo, R.C.H. & N.J. Thomas 2019. Updated Inventory of Amphibians, Reptiles and Mammals of the Bukit Timah Nature Reserve, Singapore. Gardens’ Bulletin Singapore 71(Suppl. 1):145-183; doi: 10.26492/gbs71(suppl.1).2019-06 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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