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Cyrtodactylus battalensis KHAN, 1993

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Reticulate Plump-bodied gecko 
SynonymCyrtodactylus battalensis KHAN 1993
Tenuidactylus battalensis — DAS 1994
Cyrtopodion (Tenuidactylus) battalensis — RÖSLER 2000: 68
Cyrtodactylus battalensis — KHAN 2003
Siwaligekko battalensis — KHAN 2003
Cyrtopodion battalense — FROST 2007 (pers. comm.)
Cyrtodactylus battalensis — BAUER et al. 2013
Cyrtodactylus battalensis — AGARWAL et al. 2014
Swaligekko battalensis — ALTAF et al. 2021 (in error) 
DistributionPakistan (Mansehra)

Type locality: Karakoram Highway, near Batgram Town, District Mansehra, NW Frontier Province, Pakistan (34° 40’ N, 73° 03’ E), elevation 800 m.  
Reproductionoviparous 
TypesHolotype: BMNH 1990.2 
DiagnosisDiagnosis: A large species of Cyrtodactylus geckos, with dorsal granular scales of body interspersed with scattered obtusely keeled flattish "tubercles"; 50-52 scales across midabdomen; 199-205 scales along the midventrum of body, from postmental to anterior of vent; 9-10 preanal pores arranged in an arch, no femoral pores; tail without marked segmentation, with two rows of small rounded caudal keelless tubercles well marked in anterior half of the tail, on each dorsolateral side; subcaudals small in several rows. The new species Cyrtodactylus battalensis (Fig. 1) is closest in body form and pholidosis to the circum-Himalayan stock of southeast Asian cyrtodactylid geckos; C. tibetinus, C. mintoni and C. dattanensis. However, C. battalensis has relatively longer body and tail, a wider range of supralabials, interorbitals, scales across midabdomen and along body midventrum (Table 1). The dorsal pattern of body has very irregular dark transverse bands, narrower than the interspaces, forming a reticulation by the union of irregular borders of the bands with each other. However, C. dattanensis comes closer to the new species is number of interorbitals, and scales the midabdomen. C. dattanensis and C. tibetinus both have a banded dorsal pattern, but the bands are broader than the interspaces, while in C. mintoni they are much narrower than the interspaces as in C. battalensis but are broken in spots on the sides instead of forming a reticulum. (Khan 1993) 
CommentSynonymy: T. batalensis [sic] in KHAN 1993: 220 is actually Cyrtopodion (T.) baturensis fide RÖSLER 2000.

Type Species: Cyrtodactylus battalensis KHAN 1993 is the type species of the genus Siwaligekko Khan 2003. For a comparison of characters among Siwaligekko, Altigekko, Indogekko, and Cyrtopodion, see Khan 2009.

Abundance: only known from the type locality (Meiri et al. 2017). 
EtymologyNamed after the Battal mountain Range in which the type locality is situated. 
References
  • Altaf, M. et al. 2021. Anthropogenic impacts on the diversity and distribution of amphibians and reptiles in the vicinity of Dhirkot, Azad Jammu and Kashmir, Pakistan. Journal of Wildlife and Ecology 5: 38-46 - get paper here
  • Baig, K. J. 1998. A new species of Tenuidactylus (Sauria: Gekkonidae) from Balochistan, Pakistan. Hamadryad 23 (2): 127-132. - get paper here
  • BAUER, AARON M.; RAFAQAT MASROOR, JAMES TITUS-MCQUILLAN, MATTHEW P. HEINICKE,, JUAN D. DAZA & TODD R. JACKMAN 2013. A preliminary phylogeny of the Palearctic naked-toed geckos (Reptilia: Squamata: Gekkonidae) with taxonomic implications. Zootaxa 3599 (4): 301–324 - get paper here
  • Das, I. 1994. The reptiles of South Asia: checklist and distributional summary. Hamadryad 19: 15-40 - get paper here
  • Hamid, H. N., Rais, M., Arif, M., & Noor, R. 2020. Amphibians and Reptiles of Sheikh Baddin National Park, Khyber Pakhtunkhwa: Diversity, Threats and Conservation Prospects. Pakistan J. Zool. 53(2): 1-4 - get paper here
  • Khan M S 1993. A new sandstone gecko from Fort Munro, Dera Ghazi Khan District, Punjab, Pakistan. Pakistan Journal of Zoology 25 (3) 1993: 217-221 - get paper here
  • Khan M S. 1993. A new angular-toed gecko from Pakistan, with remarks on the taxonomy and a key to the species belonging to genus Cyrtodactylus (Reptilia: Sauria: Gekkonidae). Pakistan Journal of Zoology 25 (1): 67-73. - get paper here
  • Khan, M.S. 2003. CHECKLIST AND KEY TO THE LIZARDS OF PAKISTAN. Pakistan J. Zool. Suppl. Ser. (1): 1-25 - get paper here
  • Khan, M.S. 2003. Anmerkungen zur Morphologie, Verbreitung und den Habitatpräferenzen einiger pakistanischer Geckos. Sauria 25 (3): 35-47 [erratum in 25 (4): 27] - get paper here
  • Khan, M.S. 2003. Questions of generic designation of angular-toed geckos of Pakistan with descriptions of three new genera (Reptilia: Gekkonidae). J. nat. hist. Wildl. 2 (2): 1-9
  • Khan, M.S. 2009. Intergeneric Relations of the Angular - Toed Geckos of Circum Western Himalayas (Sauria: Gekkonidae). Pakistan J. Zool., vol. 41(1), Fjp. 29-34 - get paper here
  • Khan, Muhammad Sharif 2005. An Overview of the Angular-toed Geckos of Pakistan (Squamata: Gekkonidae). Gekko 4 (2): 20-30
  • Krysko, K. L.; Rehman, H. & Auffenberg, K. 2007. A new species of Cyrtopodion (Gekkonidae: Gekkoninae) from Pakistan. Herpetologica 63 (1): 100-113 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
 
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