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Cyrtodactylus langkawiensis GRISMER, WOOD, QUAH, ANUAR, MUIN, SUMONTHA, AHMAD, BAUER, WANGKULANGKUL, GRISMER & PAUWELS, 2012

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Langkawi Island Bent-toed Gecko 
SynonymCyrtodactylus langkawiensis GRISMER, WOOD, QUAH, ANUAR, MUIN, SUMONTHA, AHMAD, BAUER, WANGKULANGKUL, GRISMER & PAUWELS 2012
Cyrtodactylus langkawiensis — GRISMER & QUAH 2019 
DistributionPeninsular Malaysia (Pulau Langkawi)

Type locality: Wat Wanaram, Pulau Langkawi, Kedah, Peninsular Malaysia (06°20.275 N, 99°52.507 E) at 35 meters elevation.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZRC 2.6966, adult male, collected on 11 October 2008 by L. L. Grismer, Norhayati A., Chan, K. O., D. Belabut, and J. L. Grismer. Paratypes.—Adult females (ZRC LSUHC 2.6967–68 and LSUHC 9123–24) collected from same locality on the same date as the holotype and by the same collectors; adult female (ZRC 2.6969) collected at same locality as holotype by Chan, K. O., C. Johnson, and L. Grismer on 19 September 2009. 
DiagnosisDiagnosis: Adult males reaching 93.3 mm SVL, adult females reaching 99.8 mm SVL; 9–12 supralabials, 8–10 infralabials; tubercles of dorsum small to moderately large with no intervening smaller tubercles; no tubercles on ventral surfaces of forelimbs, gular region, or in ventrolateral body fold; 34–44 paravertebral tubercles; 21–25 longitudinal rows of dorsal tubercles; 38–43 rows of ventral scales; 19–21 subdigital lamellae on fourth toe; 30 femoro-precloacal pores in males; dorsum not bearing a scattered pattern of white tubercles; four rarely five dark body bands in adults lacking lightened centers and light colored tubercles; band to interspace ratio 0.75–1.00; 11–16 dark caudal bands on original tail; white caudal bands in adults infused with dark pigmentation; and posterior portion of tail in hatchlings and juveniles white. These characters are scored across all species of the Cyrtodactylus pulchellus complex in Table 6 [GRISMER et al. 2012]. 
CommentAbundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet, langkawiensis, refers to Langkawi Island to where this species was first discovered and may be endemic. 
References
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L.; P. L. WOOD, JR., SHAHRUL ANUAR, H. R. DAVIS, A. J. COBOS & M. L. MURDOCH 2016. A new species of karst forest Bent-toed Gecko (genus Cyrtodactylus Gray) not yet threatened by foreign cement companies and a summary of Peninsular Malaysia’s endemic karst forest herpetofauna and the need for its conservation. Zootaxa 4061 (1): 001–017 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR, SHAHRUL ANUAR, MARTA S. GRISMER, EVAN S. H. QUAH, MATTHEW L. MURDOCH, MOHD ABDUL MUIN, HAYDEN R. DAVIS, CÉSAR AGUILAR, RANDY KLABACKA, ANTHONY J. COBOS, ANCHALEE AOWPHOL, JACK W. SITES, JR 2016. Two new Bent-toed Geckos of the Cyrtodactylus pulchellus complex from Peninsular Malaysia and multiple instances of convergent adaptation to limestone forest ecosystems. Zootaxa 4105 (5): 401-429 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR., EVAN S. H. QUAH, SHAHRUL ANUAR, MOHD. ABDUL MUIN, MONTRI SUMONTHA, NORHAYATI AHMAD, AARON M. BAUER, SANSAREEYA WANGKULANGKUL, JESSE L. GRISMER & OLIVIER S. G. PAUWELS 2012. A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa 3520: 1–55
  • Grismer, L.L.; PERRY L. WOOD, SHAHRUL ANUAR, EVAN S. H. QUAH, MOHD ABDUL MUIN, MAKETAB MOHAMED, CHAN KIN ONN, ALEXANDRA X. SUMARLI, ARIEL I. LOREDO, HEATHER M. HEINZ 2014. The phylogenetic relationships of three new species of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from poorly explored regions in northeastern Peninsular Malaysia. Zootaxa 3786 (3): 359–381 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • QUAH, EVAN S. H.; L. LEE GRISMER, PERRY L. WOOD, JR. & SHAHRUL ANUAR MOHD SAH 2019. The discovery and description of a new species of Bent-toed Gecko of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa 4668 (1): 051–075 - get paper here
  • Termprayoon, K., Rujirawan, A., Grismer, L. L., Wood Jr, P. L., & Aowphol, A. 2021. Taxonomic reassessment and phylogenetic placement of Cyrtodactylus phuketensis (Reptilia, Gekkonidae) based on morphological and molecular evidence. ZooKeys, 1040, 91 - get paper here
  • WOOD JR, PERRY L.; L. LEE GRISMER, MOHD ABDUL MUIN, SHAHRUL ANUAR, JAMIE R. OAKS 2020. A new potentially endangered limestone-associated Bent-toed Gecko of the Cyrtodactylus pulchellus (Squamata: Gekkonidae) complex from northern Peninsular Malaysia. Zootaxa 4751 (3): 437–460 - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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