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Cyrtodactylus atremus KRAUS & WEIJOLA, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common Names 
SynonymCyrtodactylus atremus KRAUS & WEIJOLA 2019
Cyrtodactylus sp. ‘Karkar’ — TALLOWIN et al. 2018 
DistributionPapua New Guinea (Madang)

Type locality: above Mom, 4.6248°S, 145.9378°E, 500–645 m a.s.l., Karkar Island, Madang Province, Papua New Guinea  
TypesHolotype: UMMZ 247747 (field tag FK 17685), mature male collected by V. Weijola, 26 February 2018.
Paratypes (n = 8). Male: Papua New Guinea: Madang Province: Karkar Island: same data as holotype (UMMZ 247746); Females: Papua New Guinea: Madang Province: Karkar Island: same data as holotype (UMMZ 247748– 51), Mom, 4.61°S, 145.92°E, 290 m a.s.l. (AMS R24744, R28901), Kevasob, 4.7°S, 145.9°E, 130 m a.s.l. (AMS R124604). 
DiagnosisDiagnosis: A moderately large (adult SVL = 108–118 mm) species of Cyrtodactylus with a small patch of somewhat enlarged precloacal scales (ca. 2–3 times size of nearby femoral scales), no enlarged femoral scales, no precloacal/femoral pores in males, and having a highly tuberculate dorsum, 22–24 rows of dorsal tubercles between the lateral skin folds, dorsal tubercles 3–5 times size of adjacent granules, 24–31 scale rows below the fourth toe, 9–11 supralabials to the center of eye, 10–13 infralabials to rictus, 40–47 mid-belly scale rows, 5–6 dark dorsal crossbands to the base of the tail, uniformly brown top of head, and lips dirty white with some brown stippling. 
EtymologyThe name is a Latinized masculine compound adjective formed from the Greek a, meaning without, and trema, meaning hole. It refers to the lack of precloacal / femoral pores in males of this species. 
  • Grismer, L. L., Wood, P. L., Poyarkov, N. A., Le, M. D., Kraus, F., Agarwal, I., ... & Grismer, J. L. 2021. Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology 71: 101–154 - get paper here
  • KRAUS, FRED; VALTER WEIJOLA 2019. New species of Cyrtodactylus (Squamata: Gekkonidae) from Karkar Island, Papua New Guinea. Zootaxa 4695 (6): 529–540 - get paper here
  • Tallowin, O. J. S., Tamar, K., Meiri, S., Allison, A., Kraus, F., Richards, S. J., & Oliver, P. M. 2018. Early insularity and subsequent mountain uplift were complementary drivers of diversification in a Melanesian lizard radiation (Gekkonidae: Cyrtodactylus). Molecular Phylogenetics and Evolution 125: 29–39 - get paper here
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