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Cyrtodactylus batucolus GRISMER, CHAN, GRISMER, WOOD & BELABUT, 2008

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Common NamesBesar Island Bent-toed Gecko 
SynonymCyrtodactylus batucolus GRISMER, CHAN, GRISMER, WOOD & BELABUT 2008
Cyrtodactylus batucolus — GRISMER 2011 
DistributionW Malaysia (Pulau Besar, Melaka)

Type locality: Panti Putera (02°06.581 N, 102°19.757 E; 39 m elevation), Pulau Besar, Melaka, Peninsular Malaysia.  
TypesHolotype: ZRC 2.6743, adult male, collected by Jesse L. Grismer, Chan K. Onn, Perry L. Wood, Jr., and L. Lee Grismer on 5 June 2008. 
DiagnosisDiagnosis. Cyrtodactylus batucolus is distinguished from all other Sunda Shelf species by having a maximum SVL of 75.2 mm; moderately sized, conical, keeled, body tubercles; tubercles occurring on occiput, forelimbs, hind limbs, and beyond base of tail; 38–42 ventral scales; no transversely enlarged, median, subcaudal scales; proximal subdigital lamellae transversely expanded; 17–19 subdigital lamellae on fourth toe; abrupt transition between postfemoral and ventral femoral scales; enlarged femoral and precloacal scales with a continuous series of 43–46 pore-bearing scales in males; precloacal groove absent, precloacal depression present; no white reticulum on head; dark blotches on body. These characters are summarized across all Sunda Shelf species in Table 1 in GRISMER et al. 2008. 
CommentAbundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet batucolus comes from the Malaysian word batu meaning rock and the Latin suffix colo meaning to inhabit or to dwell in. 
  • Chan Kin Onn & Norhayati Ahmad 2010. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa 2389: 47–56 - get paper here
  • Goldberg, S. R. and L. L. Grismer 2016. Cyrtodactylus batucolus (Besar Island Bent-toed Gecko) reproduction. Herpetological Review 47(3): 464.
  • Grismer, L. Lee & Ahmad, N. 2008. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa 1924: 53–68 - get paper here
  • Grismer, L. Lee; Chan K. Onn; Grismer, J.K.; Wood, P.L. & Belabut, D. 2008. Three new species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia. Zootaxa 1921: 1–23 - get paper here
  • Grismer, L.; Lee Perry L. Wood, Jr. and Kelvin K. P. Lim 2012. Cyrtodactylus majulah, a new species of bent-toed gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology 60 (2): 487-499 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Onn, C.K.; Griserm, L.L.; Wood, P.L.; Grismer, J.L. & Ahmad, N. 2009. Preliminary Checklist of the Herpetofauna of Pulau Besar, Melaka, Malaysia. Tropical Life Sciences Research, 20(1): 81–87
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