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Cyrtodactylus laangensis MURDOCH, GRISMER, WOOD, NEANG, POYARKOV, TRI, NAZAROV, AOWPHOL, PAUWELS, NGUYEN & GRISMER, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Phnom Laang Bent-toed Gecko 
SynonymCyrtodactylus laangensis MURDOCH, GRISMER, WOOD, NEANG, POYARKOV, TRI, NAZAROV, AOWPHOL, PAUWELS, NGUYEN & GRISMER 2019: 42 
DistributionCambodia (Kampot: Phnom Laang)

Type locality: Phnom Laang, Kampot Province, Cambodia (10 42’13’’N, 104 20’57’’ E).  
Reproduction 
TypesHolotype: LSUHC 8773, adult male, collected on 15 December 2007 by Neang Thy, L. Lee Grismer, and Jeremy Holden. Paratypes. Adult males LSUHC 8771–72, adult females LSUHC 8770, 8774, and T 4653 bear the same collection data as the holotype. 
DiagnosisDiagnosis. Adult males reaching 76.9 mm SVL, adult females reaching 82.2 mm SVL; 7–9 supralabials, 10 or 11 infralabials; 29–32 paravertebral tubercles; 17 or 18 longitudinal rows of dorsal tubercles; 37–40 rows of ventral scales; seven expanded subdigital lamellae proximal to the digital inflection, 11–13 unmodified, distal, subdigital lamellae; 18–20 total subdigital lamellae on fourth toe; enlarged femoral scales either absent or separated from precloacal scales by a diastema of smaller scales; when present 13–16 enlarged femoral scales with proximal enlarged femoral scales less than half the size of distal femoral scales; 7–9 enlarged precloacal scales with pores on each in males; two or three rows of enlarged post-precloacal scales; 2–4 postcloacal tubercles; no interdigital pocketing; dark pigmented blotches on top of head present or absent; posterior border of nuchal loop rounded; and four dark body bands (summarized in Tables 7 + 13 in Murdoch et al. 2019).

Comparisons. Cyrtodactylus laangensis sp. nov. is a member of the eastern group and the sister species to Cyrtodactylus bokorensis sp. nov. from the Bokor Plateau, Kampot Province, Cambodia from which it is separated by 3.9% sequence divergence (Table 4 in Murdoch et al. 2019). The PCA and DAPC analyses indicate that Cyrtodactylus laangensis sp. nov. is very isolated in morphospace from all other species within the complex (Fig. 6, 7). Cyrtodactylus laangensis sp. nov. is well-differentiated from all other members of the C. intermedius complex by having varying combinations of statistically different mean values of supralabial and infralabial scales, paravertebral tubercles, ventral scales, expanded, and total number of subdigital lamellae, enlarged femoral scales, precloacal scales, and postcloacal tubercles (Table 6). It differs from C. bokorensis by having a rounded posterior border of the nuchal loop as opposed to a pointed border. It is differentiated from all other members of the C. intermedius complex in having a diastema between the enlarged femoral scales and the precloacal scales, with the exception of C. bokorensis for which contact is variable. It is further differentiated from C. cardamomensis and C. thylacodactylus in that the proximal most scales of its short femoral scale row are less than half the size of the distal scales. It is further differentiated from C. thylacodactylus in lacking interdigital pocketing (Table 7).
 
CommentHabitat: karst hills surrounded by highly disturbed scrub vegetation but the hill itself bears typical ultramafic vegetation on its cliff-faces. Some specimens were found crawling on both large boulders and small rocks and others were found as high as 5 m above the ground on (cave-like) chamber walls.

Despite being separated by only 25km from its sister species, C. bokorensis, the two species have an uncorrected sequence divergence of 3.9% and the morphological and morphometric differences between the two species are larger than between species having greater sequence divergence (Table 4, Fig. 5 in Murdoch et al. 2019). 
EtymologyThe specific epithet, laangensis, is an adjective in reference to type locality of the karst formation Phnom Laang to which it is presumably endemic. 
References
  • MURDOCH, MATTHEW L.; L. LEE GRISMER, PERRY L. JR. WOOD, THY NEANG, NIKOLAY A. POYARKOV, NGO VAN TRI, ROMAN A. NAZAROV, ANCHALEE AOWPHOL, OLIVIER S.G. PAUWELS, HUNG NGOC NGUYEN, JESSE L. GRISMER 2019. Six new species of the Cyrtodactylus intermedius complex (Squamata: Gekkonidae) from the Cardamom Mountains and associated highlands of Southeast Asia. Zootaxa 4554 (1): 001–062 - get paper here
 
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