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Cyrtodactylus pinlaungensis GRISMER, WOOD, QUAH, THURA, OAKS & LIN, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesPinlaung Bent-toed Gecko 
SynonymCyrtodactylus pinlaungensis GRISMER, WOOD, QUAH, THURA, OAKS & LIN 2019 
DistributionMyanmar (Pa-O District, Shan State)

Type locality: karstic hills immediately behind the Pin-laung Hotel, Pinlaung Township, Pa-O District, Shan State, Myanmar (20.13042°N, 96.78620°E; 1534 m in elevation).  
Reproduction 
TypesHolotype: LSUHC 14280, Adult male, collected on 12 November 2018 at 2000 hrs by Evan S. H. Quah, Myint Kyaw Thura, Jamie R. Oaks, Perry L. Wood Jr., Aung Lin and L. Lee Grismer.
Paratypes: Adult females LSUHC 14278–79 bear the same collection data as the holotype. Adult male LSUHC 14307, adult female LSUHC 14306, and subadult female 14308 were collected on 13 November 2018 between 21:00–23:30 hrs by Myint Kyaw Thura, Jamie R. Oaks, Evan S. H. Quah, Perry L. Wood Jr., Aung Lin and L. Lee Grismer at Wingabar Cave, Pinlaung Township, Pa-O District, Shan State, Myanmar (20.06936°N, 96.76998°E; 1464 m in elevation). 
DiagnosisDiagnosis. Cyrtodactylus pinlaungensis sp. nov. differs from all species in the C. linnwayensis group by having the combination of seven or eight supralabials; 8–10 infralabials; 36–42 paravertebral tubercles; 24–29 longitudi- nal rows of dorsal tubercles; 35–43 ventral scales; 5–7 pore-bearing precloacal scales in males; 9–14 pore-bearing femoral scales in males; 6–8 enlarged precloacal scales; 4–6 enlarged post-precloacal scales; 24–31 enlarged femo- ral scales; eight or nine transversely expanded subdigital lamellae on the fourth toe; 12–15 unexpanded subdigital lamellae on the fourth toe; 20–23 total subdigital lamellae on the fourth toe; raised and strongly keeled dorsal tubercles that extend beyond the postcloacal swelling; four or five regularly shaped dorsal bands bearing lightened centers; and a mature regenerated tail with a faint to prominent spotted pattern (Table 4 in Grismer et al. 2019).

Comparisons: (summarized in Figs. 3, 7, Table 4 in Grismer et al. 2019). Cyrtodactylus pinlaungensis sp. nov. differs from all other species in the C. linnwayensis group in having a significantly higher mean number of paravertebral tubercles (38.8 versus 28.6–33.6, collectively) and dorsal tubercles that extend beyond the postcloacal swelling as opposed to terminating anterior to it. Cyrtodactylus pinlaungensis sp. nov. differs from C. linnwayensis and C. shwetaungorum in having a higher mean number of longitudinal rows of tubercles (26.2 versus 17.8 and 19.0, respectively), a significantly lower mean number of enlarged precloacal scales (7.2 versus 10.1 and 9.8, respectively), and raised and strongly keeled versus low and weekly keeled dorsal tubercles. It differs further from C. ywanganensis in having a significantly higher mean number of post-precloacal scales (5.0 versus 3.0) and ventral scales (39.3 versus 35.0), and regularly shaped dorsal bands bearing lightened centers as opposed to jagged dorsal bands lacking lightened centers. Cyrtodactylus pinlaungensis sp. nov. differs further from C. shwetaungorum in having a significantly lower mean number of post- precloacal scales (5.0 versus 3.2 and 3.0, respectively) and a mature regenerated tail bearing as opposed to lacking dark spots. Cyrtodactylus pinlaungensis sp. nov. is completely separated from all species in the C. linnwayensis group in the PCA and DAPC. Given the small sample size for C. pinlaungensis sp. nov. (N=6) and C. ywanganensis (N=5), the statistical significance of some of these characters may increase or decrease with the acquisition of ad- ditional specimens. 
Comment 
EtymologyNamed after the Pin-laung Hotel where the type was found. 
References
  • GRISMER, L. LEE; PERRY L. WOOD JR., EVAN S. H. QUAH, MYINT KYAW THURA,<br />JAMIE R. OAKS & AUNG LIN 2019. A new species of Bent-toed Gecko (Squamata, Gekkonidae, Cyrtodactylus) from the Shan Plateau in eastern Myanmar (Burma). Zootaxa 4624 (3): 301–321 - get paper here
 
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