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Cyrtodactylus pulchellus GRAY, 1827

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Malayan Forest Gecko, Banded Bent-toed Gecko
G: Malaiischer Bogenfingergecko 
SynonymCyrtodactylus pulchella GRAY 1827: 56
Cyrtodactylus puchellus — HARDWICKE & GRAY 1835 (in error)
Gymnodactylus pulchellus — DUMÉRIL & BIBRON 1836: 423
Gymnodactylus pulchellus — STOLICZKA 1873
Gymnodactylus pulchellus — BOULENGER 1885: 46
Cyrtodactylus pulchellus — TAYLOR 1963: 714
Cyrtodactylus pulchellus — KAHL et al. 1980: 139
Cyrtodactylus pulchellus — MANTHEY & GROSSMANN 1997: 226
Cyrtodactylus pulchellus — COX et al. 1998: 86
Cyrtodactylus (Cyrtodactylus) pulchellus — RÖSLER 2000: 67
Cyrtodactylus abrae WELLS 2002 (fide Glen Shea, pers. comm. 19 March 2013)
Cyrtodactylus pulchellus — GRISMER 2011 
DistributionMalaysian Peninsula (Penang Island), Thailand

Type locality: “India”; corrected to “Singapore” by GRAY 1845. Restricted to Penang Island by Grismer 2012.  
Reproductionoviparous 
TypesHolotype: BMNH 
CommentDistribution: A report from Myanmar (Das and Lim 2000), remains to be verified. Reports from India are in error (A. Bauer, pers. comm. 30 Aug 2011).

Type species: Cyrtodactylus pulchella GRAY 1827 is the type species of the genus Cyrtodactylus GRAY 1827.

A diagnostis character of Cyrtodactylus is their possession of vertical pupils and bent, padless toes.

Diagnosis.– Adult males reaching 114.1 mm SVL, adult females reaching 113.3 mm SVL; 9–11 supralabials, 8–10 infralabials; tubercles of dorsum moderately large, strongly keeled and widely separated with no intervening smaller tubercles; no tubercles on ventral surface of forelimbs, gular region, or in ventrolateral body fold or these regions may be weekly tuberculated; 33–43 paravertebral tubercles; 22–26 longitudinal rows of dorsal tubercles; 29–34 rows of ventral scales; 21–26 subdigital lamellae on fourth toe; 33–39 femoro-precloacal pores in males; dorsum not bearing scattered, white tubercles; four dark body bands with a body band/interspace ratio of 0.75–1.25; 8–10 dark caudal bands on original tail; white caudal bands in adults immaculate; and posterior portion of tail hatchlings and juveniles not white (Table 4). These characters are scored across all species of the Cyrtodactylus pulchellus complex in Table 6 [GRISMER et al. 2012].

Synonymy: Cyrtodactylus abrae WELLS 2002 is a nomen nudum fide COUPER & AMEY 2004. 
EtymologyNamed after Latin “pulchellus” = pretty. 
References
  • Bauer, A.M. 2013. Geckos - The Animal Answer Guide. Johns Hopkins University Press, 159 pp.
  • Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Chan Kin Onn & Norhayati Ahmad 2010. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa 2389: 47–56 - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Couper, P. J.; Amey, A. 2004. Comments on the taxonomic status of Cyrtodactylus abrae WELLS 2002. Memoirs of the Queensland Museum 49(2): 648.
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  • Das, I. & Lim, L.J. 2000. A new species of Cyrtodactylus (Sauria: Gekkonidae) from Pulau Tioman. Raffles Bull. Zool. (Singapore) 48 (2): 223-231 - get paper here
  • Duméril, A.M. C. and G. Bibron. 1836. Erpetologie Générale ou Histoire Naturelle Complete des Reptiles. Vol.3. Libr. Encyclopédique Roret, Paris, 528 pp. - get paper here
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  • GRISMER, L. LEE; PERRY L. WOOD, JR, SHAHRUL ANUAR, MARTA S. GRISMER, EVAN S. H. QUAH, MATTHEW L. MURDOCH, MOHD ABDUL MUIN, HAYDEN R. DAVIS, CÉSAR AGUILAR, RANDY KLABACKA, ANTHONY J. COBOS, ANCHALEE AOWPHOL, JACK W. SITES, JR 2016. Two new Bent-toed Geckos of the Cyrtodactylus pulchellus complex from Peninsular Malaysia and multiple instances of convergent adaptation to limestone forest ecosystems. Zootaxa 4105 (5): 401-429 - get paper here
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  • Rösler, H. 2000. Die postanale Beschuppung bei Cyrtodactylus GRAY 1827 und Cyrtopodion FITZINGER 1843 - funktionelle und taxonomische Aspekte (Sauria: Gekkonidae). Gekkota 2: 154-207
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