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Cyrtodactylus pulchellus GRAY, 1827

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Higher TaxaGekkonidae, Sauria (lizards: geckos) 
Subspecies 
Common NamesE: Malayan Forest Gecko, Banded Bent-toed Gecko
G: Malaiischer Bogenfingergecko 
SynonymCyrtodactylus pulchella GRAY 1827: 56
Cyrtodactylus puchellus — HARDWICKE & GRAY 1835 (in error)
Gymnodactylus pulchellus — DUMÉRIL & BIBRON 1836: 423
Gymnodactylus pulchellus — STOLICZKA 1873
Gymnodactylus pulchellus — BOULENGER 1885: 46
Cyrtodactylus pulchellus — TAYLOR 1963: 714
Cyrtodactylus pulchellus — KAHL et al. 1980: 139
Cyrtodactylus pulchellus — MANTHEY & GROSSMANN 1997: 226
Cyrtodactylus pulchellus — COX et al. 1998: 86
Cyrtodactylus (Cyrtodactylus) pulchellus — RÖSLER 2000: 67
Cyrtodactylus abrae WELLS 2002 (fide Glen Shea, pers. comm. 19 March 2013)
Cyrtodactylus pulchellus — GRISMER 2011 
DistributionMalaysian Peninsula (Penang Island)

Type locality: “India”; corrected to “Singapore” by GRAY 1845. Restricted to Penang Island by Grismer 2012. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
TypesType: BMNH (and possibly additional specimens). 
CommentDistribution: A report from Myanmar (Das and Lim 2000), remains to be verified. Reports from India are in error (A. Bauer, pers. comm. 30 Aug 2011).

Type species: Cyrtodactylus pulchella GRAY 1827 is the type species of the genus Cyrtodactylus GRAY 1827.

A diagnostis character of Cyrtodactylus is their possession of vertical pupils and bent, padless toes.

Diagnosis.– Adult males reaching 114.1 mm SVL, adult females reaching 113.3 mm SVL; 9–11 supralabials, 8–10 infralabials; tubercles of dorsum moderately large, strongly keeled and widely separated with no intervening smaller tubercles; no tubercles on ventral surface of forelimbs, gular region, or in ventrolateral body fold or these regions may be weekly tuberculated; 33–43 paravertebral tubercles; 22–26 longitudinal rows of dorsal tubercles; 29–34 rows of ventral scales; 21–26 subdigital lamellae on fourth toe; 33–39 femoro-precloacal pores in males; dorsum not bearing scattered, white tubercles; four dark body bands with a body band/interspace ratio of 0.75–1.25; 8–10 dark caudal bands on original tail; white caudal bands in adults immaculate; and posterior portion of tail hatchlings and juveniles not white (Table 4). These characters are scored across all species of the Cyrtodactylus pulchellus complex in Table 6 [GRISMER et al. 2012].

Synonymy: Cyrtodactylus abrae WELLS 2002 is a nomen nudum fide COUPER & AMEY 2004. 
EtymologyEtymology: Named after Latin “pulchellus” = pretty. 
References
  • Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Chan Kin Onn & Norhayati Ahmad 2010. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from northeastern Peninsular Malaysia, Malaysia. Zootaxa 2389: 47–56 - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Couper, P. J.; Amey, A. 2004. Comments on the taxonomic status of Cyrtodactylus abrae WELLS 2002. Memoirs of the Queensland Museum 49(2): 648.
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Das, I. & Lim, L.J. 2000. A new species of Cyrtodactylus (Sauria: Gekkonidae) from Pulau Tioman. Raffles Bull. Zool. (Singapore) 48 (2): 223-231
  • Duméril, A.M. C. and G. Bibron. 1836. Erpetologie Générale ou Histoire Naturelle Complete des Reptiles. Vol.3. Libr. Encyclopédique Roret, Paris, 528 pp. - get paper here
  • Gray, J. E. 1827. A Synopsis of the Genera of Saurian Reptiles in which some new Genera are indicated, and the others reviewed by actual Examination. Philos. Mag., London, 2 (2): 54-58.
  • Gray, J. E. 1845. Catalogue of the specimens of lizards in the collection of the British Museum. Trustees of die British Museum/Edward Newman, London: xxvii + 289 pp. - get paper here
  • Grismer L. Lee 2005. New species of Bent-toed Gecko (Cyrtodactylus Gray 1827) from Pulau Aur, Johor, West Malaysia. Journal of Herpetology 39 (3):424-432. - get paper here
  • Grismer L. Lee & Leong, T.M. 2005. New Species of Cyrtodactylus (Squamata: Gekkonidae) from Southern Peninsular Malaysia. Journal of Herpetology 39 (4): 584-591 - get paper here
  • Grismer, L. Lee & Ahmad, N. 2008. A new insular species of Cyrtodactylus (Squamata: Gekkonidae) from the Langkawi Archipelago, Kedah, Peninsular Malaysia. Zootaxa 1924: 53–68 - get paper here
  • Grismer, L. Lee; Chan K. Onn, Jesse L. Grismer, Perry L. Wood, Jr., and A. Norhayati 2010. A CHECKLIST OF THE HERPETOFAUNA OF THE BANJARAN BINTANG, PENINSULAR MALAYSIA. Russian Journal of Herpetology 17 (2): 147-160 - get paper here
  • Grismer, L. Lee; Chan K. Onn; Grismer, J.K.; Wood, P.L. & Belabut, D. 2008. Three new species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia. Zootaxa 1921: 1–23 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, SHAHRUL ANUAR, EVAN S. H. QUAH, MOHD ABDUL MUIN, MAKETAB MOHAMED, CH 2014. The phylogenetic relationships of three new species of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from poorly explored regions in northeastern Peninsular Malaysia Zootaxa 3786 (3): 359–381 - get paper here
  • Grismer, L.; Lee Perry L. Wood, Jr. and Kelvin K. P. Lim 2012. Cyrtodactylus majulah, a new species of bent-toed gecko (Reptilia: Squamata: Gekkonidae) from Singapore and the Riau Archipelago. The Raffles Bulletin of Zoology 60 (2): 487-499 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155]
  • Kahl B, Gaupp P & Schmidt G 1980. Das Terrarium. Falken Verlag, Niederhausen (Germany), 336 pp.
  • Krysko, K.L.; Rehman, H. & Auffenberg, K. 2007. A new species of Cyrtopodion (Gekkonidae: Gekkoninae) from Pakistan. Herpetologica 63 (1): 100-113 - get paper here
  • Linkem, Charles W.; Jimmy A. McGuire, Christopher J. Hayden, Mohammed Iqbal Setiadi, David P. Bickfo 2008. A New Species of Bent-Toe Gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica 64 (2): 224-234 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Pauwels,O.S.G. et al. 2000. Herpetological investigations in Phang-Nga Province, southern Peninsular Thailand, with a list of reptile species and notes on their biology. Dumerilia 4 (2): 123-154 - get paper here
  • Quang, Hoang Xuan, Nikolai L. Orlov, Natalia B. Ananjeva, Andrew G. Johns, Hoang N. Thao and Dau Q. 2007. Description of a new species of the genus Cyrtodactylus Gray, 1827(Squamata: Sauria: Gekkonidae) from the Karst of north central Vietnam. Russian Journal of Herpetology 14 (2):98-106 - get paper here
  • Rösler, H. 2000. Die postanale Beschuppung bei Cyrtodactylus GRAY 1827 und Cyrtopodion FITZINGER 1843 - funktionelle und taxonomische Aspekte (Sauria: Gekkonidae). Gekkota 2: 154-207
  • Rösler, H. & Glaw, F. 2008. A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus. Zootaxa 1729: 8–22 - get paper here
  • SHEA, GLENN; PATRICK COUPER, JESSICA WORTHINGTON WILMER & ANDREW AMEY 2011. Revision of the genus Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Australia. Zootaxa 3146: 1–63
  • Stoliczka,F. 1873. Notes on some species of Malayan Amphibia and reptilia. J. Asiat. Soc. Bengal 42: 111-126 - get paper here
  • Sumontha, M., Panitvong, N. & Deein, G. 2010. Cyrtodactylus auribalteatus (Squamata: Gekkonidae), a new cave- dwelling gecko from Phitsanulok Province, Thailand. Zootaxa 2370: 53–64 - get paper here
  • Taylor, E.H. 1963. The lizards of Thailand. Univ. Kansas Sci. Bull. 44: 687-1077. - get paper here
  • Tri, Ngo Van and L. Lee Grismer 2010. A new karst dwelling Cyrtodactylus (Squamata: Gekkonidae) from Son La Province, north-western Vietnam. Hamadryad 35 (1): 84 – 95
  • Welton, L.J, Siler, C.D., Diesmos, A.C. & Brown, R.M. 2010. Phylogeny-based species delimitation of southern Philippines bent-toed geckos and a new species of Cyrtodactylus (Squamata: Gekkonidae) from western Mindanao and the Sulu Archipelago. Zootaxa 2390: 49–68 - get paper here
  • Welton, Luke J.; Cameron D. Siler, Arvin Diesmos, and Rafe M. Brown 2009. A NEW BENT-TOED GECKO (GENUS CYRTODACTYLUS) FROM SOUTHERN PALAWAN ISLAND, PHILIPPINES AND CLARIFICATION OF THE TAXONOMIC STATUS OF C. ANNULATUS. Herpetologica 65 (3): 328-343 - get paper here
  • Youmans, Timothy M. & L. Lee Grismer 2006. A new species of Cyrtodactylus (Reptilia: Squamata: Gekkonidae) from the Seribuat Archipelago, West Malaysia. Herpetological Natural History 10 (1): 61-70
 
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