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Cyrtodactylus pyadalinensis GRISMER, WOOD, THURA, WIN & QUAH, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common Names 
SynonymCyrtodactylus pyadalinensis GRISMER, WOOD, THURA, WIN & QUAH 2019 
DistributionMyanmar (Shan)

Type locality: Panluang-Pyadalin Cave Wildlife Sanctuary, Ywangan Township, Shan State, Myanmar (21.107000°N, 96.352111°E; 220 m in elevation).  
Reproduction 
TypesHolotype: CAS 226143, subadult male collected during the evening on 16 July 2002 by G.O.U. Wogan, R. S. Lucas, J. V. Vindum, Thin Thin, and A. K. Shein.
Paratypes. Subadult male CAS 226142 collected during the evening on 16 July 2002 by Htun Win from Panluang-Pyadalin Cave Wildlife Sanctuary, Ywangan Township, Shan State, Myanmar (21.115801°N, 96.360694°E; 346 m in elevation). Adult female LSUHC 13932 collected at 2100 hrs on 26 March 2018 by Perry L. Wood Jr., Nyo Min Htwe, and L. Lee Grismer from immediately outside the Pyadalin Cave, Panluang-Pyadalin Cave Wildlife Sanctuary, Ywangan Township, Taunggyi District, Shan State, Myanmar (21.13275°N, 96.34026°E; 306m in elevation.) 
DiagnosisDiagnosis. Cyrtodactylus pyadalinensis sp. nov. differs from all other species in the peguensis group by having the unique combination of eight supralabials and 6–8 infralabials; 31–33 paravertebral tubercles; 19–21 longitudinal rows of body tubercles; 38–40 ventral scales; 16–18 subdigital lamellae on the fourth toe; 14 or 15 femoral pores in males; nine or 10 precloacal pores in males; two or three rows of enlarged, post-precloacal scales; top of head bearing dark blotches; 4–6 dark body bands; dark body bands lacking paravertebral elements; and maximum SVL of 72.1 mm (Table 4 in Grismer et al. 2019).

Comparisons. Cyrtodactylus pyadalinensis sp. nov. descends from one of the deeper divergences of the peguensis group and the sister species to the clade (C. niyniykyawi sp. nov. (C. peguensis (C. pyinyaungensis and C. myintkyawthurai))) from which it differs by an uncorrected pairwise sequence divergence of 9.0–10.3%. From C. meersi and C. annandalei which occur outside this clade, it differs by 10.7–11.0% and 14.0–14.3%, respectively. It differs from all other species except C. nyinyikyawi sp. nov. in the dark-brown dorsal bands lacking as opposed to having paravertebral elements. It differs further from C. annandalei in that the top of the head is blotched as opposed to being unicolor. Differences from C. nyinyikyawi sp. nov. are presented in the Comparisons section above. Statistically significant mean differences in meristic characters among C. pyadalinensis sp. nov., C. myintkyawthurai, and C. pyinyaungensis are presented in Tables 3 and 4 in Grismer et al. 2019. 
CommentHabitat: The habitat is composed of low-lying, highly eroded terrain and scree of the Nwalabo Mountains. It bears scattered karstic rocks and boulders surrounded by disturbed, drought-adapted, scrub Indiang Forest vegetation that is seasonally burned. All specimens were found at night between 1900 and 2300 hrs among small rocks and leaf-leaf litter. 
EtymologyThe specific epithet, pyadalinensis, is a toponym referring to the type locality in the vicinity of the Pyadalin Cave. 
References
  • GRISMER, L. LEE; PERRY L. JR. WOOD, MYINT KYAW THURA, NAY MYO WIN, EVAN S. H. QUAH 2019. Two more new species of the Cyrtodactylus peguensis group (Squamata: Gekkonidae) from the fringes of the Ayeyarwady Basin, Myanmar. Zootaxa 4577 (2): 274–294 - get paper here
 
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