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Cyrtodactylus spinosus LINKEM, MCGUIRE, HAYDEN, SETIADI, BICKFORD & BROWN, 2008

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common Names 
SynonymCyrtodactylus spinosus LINKEM, MCGUIRE, HAYDEN, SETIADI, BICKFORD & BROWN 2008
Cyrtodactylus spinosus — MECKE et al. 2016 
DistributionIndonesia (Sulawesi)

Type locality: Indonesia: Sulawesi Island: Sulawesi Tenggah Province: Kabupaten Donggala: Kecematan Kulawi: Desa Mataue: Lore-Lindu National Park (01.44883° S, 119.99483° E) at 696 m elevation.  
Reproductionoviparous 
TypesHolotype: MZB 7024 (BSI-FS 1694), adult male, collected 8 November 2004 at 19:35 h, from. Collected by CJH, RMB, JAM and CWL. 
CommentDiagnosis.—Cyrtodactylus spinosus is distinguished from all other Cyrtodactylus species by the following characters: a row of spines along ventrolateral body fold; six lateral rows of small, unkeeled body tubercles, with most ventral row intermixed with spines; two spines on temporal region of head; 31 spine-adorned annuli encircling original tail; tubercles on fore- and hind limbs; spines on postantefemoral portion of hind limb. Additional characters distinguishing this species include: proximal subdigital lamellae transversely expanded; 19–21 subdigital lamellae on toe IV; 38–44 mid body ventral scales; most scales in femoral region small, granular; 7–12 enlarged femoral series scales lacking pores; presence of pre-cloacal groove in males (absent in females); presence of pre-cloacal pores (12–13) in a chevron-shaped groove; subcaudals not transversely expanded; three chevron-shaped dark bands on a grayish-brown background. [from LINKEM et al. 2008]. 
EtymologyThe name is derived from the Latin word spina which means ‘‘spiny’’ or ‘‘thorny’’ in reference to the unique spines possessed by this species. 
References
  • GRISMER, L. LEE; SHAHRUL ANUAR, EVAN QUAH, MOHD ABDUL MUIN, CHAN KIN ONN, JESSE L. GRISMER & NORHAYATI AHMAD 2010. A new spiny, prehensile-tailed species of Cyrtodactylus (Squamata: Gekkonidae) from Peninsular Malaysia with a preliminary hypothesis of relationships based on morphology. Zootaxa 2625: 40–52 - get paper here
  • HARTMANN, LUKAS; SVEN MECKE, MAX KIECKBUSCH, FELIX MADER, HINRICH KAISER 2016. A new species of bent-toed gecko, genus Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae), from Jawa Timur Province, Java, Indonesia, with taxonomic remarks on C. fumosus (Müller, 1895). Zootaxa 4067 (5): 552–568 - get paper here
  • Iskandar, D.; Rachmansah, A. & Umilaela 2011. A new bent-toed gecko of the genus Cyrtodactylus Gray, 1827 (Reptilia, Gekkonidae) from Mount Tompotika, eastern peninsula of Sulawesi, Indonesia. Zootaxa 2838: 65–78 - get paper here
  • Koch, A. 2011. The Amphibians and Reptiles of Sulawesi: Underestimated Diversity in a Dynamic Environment. In: F.E. Zachos and J.C. Habel (eds.), Biodiversity Hotspots. Springer, Berlin, p. 383-404
  • Koch, A. 2012. Discovery, Diversity, and Distribution of the Amphibians and Reptiles of Sulawesi and its offshore islands. Edition Chimaira, 374 pp. [ISBN 978-3-89973-432-4] - get paper here
  • Linkem, Charles W.; Jimmy A. McGuire, Christopher J. Hayden, Mohammed Iqbal Setiadi, David P. Bickford, and Rafe M. Brown 2008. A New Species of Bent-Toe Gecko (Gekkonidae: Cyrtodactylus) from Sulawesi Island, Eastern Indonesia. Herpetologica 64 (2): 224-234 - get paper here
  • Mecke, Sven; Lukas Hartmann, Felix Mader, Max Kieckbusch, Hinrich Kaiser 2016. Redescription of Cyrtodactylus fumosus (Müller, 1895) (Reptilia: Squamata: Gekkonidae), with a revised identification key to the bent-toed geckos of Sulawesi Acta Herpetologica 11 (2): 151-160 - get paper here
  • MECKE, SVEN; MAX KIECKBUSCH, LUKAS HARTMANN, HINRICH KAISER 2016. Historical considerations and comments on the type series of Cyrtodactylus marmoratus Gray, 1831, with an updated comparative table for the bent-toed geckos of the Sunda Islands and Sulawesi. Zootaxa 4175 (4): 353–365
  • OLIVER , PAUL; PAUL EDGAR, MUMPUNI, DJOKO T. ISKANDAR & RON LILLEY 2009. A new species of bent-toed gecko (Cyrtodactylus: Gekkonidae) from Seram Island, Indonesia. Zootaxa 2115: 47-55 - get paper here
  • RIYANTO, AWAL; HELLEN KURNIATI, ANDREW ENGILIS, JR 2016. A new Bent-toed gecko (Squamata: Gekkonidae) from the Mekongga Mountains, South East Sulawesi, Indonesia. Zootaxa 4109 (1): 059-072 - get paper here
  • Wanger, T. C., I. Motzke, S. Saleh & D. T. Iskandar 2011. The amphibians and reptiles of the Lore Lindu National Park area, Central Sulawesi, Indonesia. Salamandra 47 (1): 17-29 - get paper here
 
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