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Cyrtodactylus wayakonei NGUYEN, KINGSADA, RÖSLER, AUER & ZIEGLER, 2010

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesChinese: 瓦氏裸趾虎 
SynonymCyrtodactylus wayakonei NGUYEN, KINGSADA, RÖSLER, AUER & ZIEGLER 2010 
DistributionN Laos (Luang Nam Tha), China (Yunnan)

Type locality: Kao Rao Cave, near Ban Nam Eng, Vieng Phoukha District, Luang Nam Tha Province, northern Laos (20°43.516’N, 101°09.239’E), elevation 730 m.  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: IEBR A.2010.01, adult male, collected on 1 May 2010 by Truong Quang Nguyen and Phouthone Kingsada. Paratypes. ZFMK 91016, adult male; MTD 47731, NUOL 2010.1, adult females; the same locality as the holotype, altitude 730–810 m a.s.l., collected between 1 and 2 May 2010 by Truong Quang Nguyen and Phouthone Kingsada (Figs. 4–5). 
DiagnosisDiagnosis: A medium sized Cyrtodactylus of up to 90 mm maximum SVL, head distinct from slender body, with flat belly. Cyrtodactylus wayakonei sp. n. can be distinguished from all congeners on the basis of the following combination of characters: 7–8 supralabials; 9–10 infralabials; rostrum bearing smooth scales; 34–45 scales between fifth supralabials; dorsal tubercles smooth to slightly keeled; 17–19 rows of dorsal tubercles; ventrolateral folds slightly developed, with large tubercles; 31–35 ventrals; 85–98 scales around midbody; 151–163 scales between mental and cloacal slit; 6–8 precloacal pores in both sexes; precloacal groove lacking; femoral scales not enlarged; no femoral pores; subcaudals somewhat enlarged, broadened; tail tubercles flat, smooth; head and dorsum with blotched to reticulated pattern. 
CommentDistribution: for a map with the type localities of Cyrtodactylus wayakonei, martini, zhenkangensis, menglianensis, phukhaensis, and caixitaoi, see Liu et al. 2023: 535 (Fig. 6). 
EtymologyThe new Cyrtodactylus species is named after Associate Professor Dr. Sengdeuane Wayakone, Director, Office of Planning and International Cooperation, National University of Laos, in recognition of his efforts to investigate and protect biodiversity in Laos. 
References
  • DAVID, PATRICK; TRUONG QUANG NGUYEN, NICOLE SCHNEIDER & THOMAS ZIEGLER 2011. A new species of the genus Cyrtodactylus Gray, 1827 from central Laos (Squamata: Gekkonidae). Zootaxa 2833: 29–40 - get paper here
  • Liu, S., Rao, D., Hou, M., Wang, Q., & Ananjeva, N. B. 2023. A New Species of Cyrtodactylus Gray, 1827 (Squamata, Gekkonidae), Previously Confused with C. wayakonei Nguyen, Kingsada, Rösler, Auer et Ziegler, 2010. Russian Journal of Herpetology, 30(6), 529-538 - get paper here
  • LUU, VINH QUANG; THOMAS CALAME, Truong QUang Nguyen, MICHAEL BONKOWSKI & THOMAS ZIEGLER 2015. A new species of Cyrtodactylus (Squamata: Gekkonidae) from the limestone forest of Khammouane Province, central Laos. Zootaxa 4058 (3): 388–402 - get paper here
  • LUU, VINH QUANG; Truong QUang Nguyen, MINH DUC LE, MICHAEL BONKOWSKI, THOMAS ZIEGLER 2016. A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa 4079 (1): 087–102 - get paper here
  • LUU, VINH QUANG;THOMAS CALAME, MICHAEL BONKOWSKI, TRUONG QUANG NGUYEN & THOMAS ZIEGLER 2014. A new species of Cyrtodactylus (Squamata: Gekkonidae) from Khammouane Province, Laos. Zootaxa 3760 (1): 054–066 - get paper here
  • NAZAROV, ROMAN; NIKOLAY A. POYARKOV, NIKOLAI L. ORLOV, TRUNG MY PHUNG, TAO THIEN NGUYEN, DUC MINH HOANG & THOMAS ZIEGLER 2012. Two new cryptic species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa 3302: 1–24 - get paper here
  • Ngo, Van Tri 2011. Cyrtodactylus martini, another new karst-dwelling Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Northwestern Vietnam. Zootaxa 2834: 33–46 - get paper here
  • NGO, VAN TRI & L. LEE GRISMER 2012. A new endemic species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Tho Chu Island, southwestern Vietnam. Zootaxa 3228: 48–60 - get paper here
  • NGUYEN, TRUONG QUANG; PHOUTHONE KINGSADA, HERBERT RÖSLER, MARKUS AUER & THOMAS ZIEGLER 2010. A new species of Cyrtodactylus (Squamata: Gekkonidae) from northern Laos. Zootaxa 2652: 1–16 - get paper here
  • SCHNEIDER, NICOLE; TRUNG MY PHUNG, MINH DUC LE,TRUONG QUANG NGUYEN, & THOMAS ZIEGLER 2014. A new Cyrtodactylus (Squamata: Gekkonidae) from Khanh Hoa Province, southern Vietnam. Zootaxa 3785 (4): 518–532 - get paper here
  • Wang K, Lyu ZT, Wang J, Qi S, Che J 2022. Updated Checklist and Zoogeographic Division of the Reptilian Fauna of Yunnan Province, China. Biodiversity Science 30 (4): 21326, 1–31 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • YUAN, Si-Qi & RAO Ding-Qi 2011. A new record of a Gekkonid (Cyrtodactylus wayakonei) from Yunnan, China. Zoological Research 32 (6): 684−688
  • Ziegler, T. 2012. Erforschung und Erhalt der Artenvielfalt: Neue Wege des Kölner Zoos in Südostasien. Zeitschrift des Kölner Zoos 55 (3): 111-
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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