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Darevskia bithynica (MÉHELY, 1909)

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Higher TaxaLacertidae, Lacertinae, Sauria, Lacertoidea, Squamata (lizards)
SubspeciesDarevskia bithynica bithynica (MÉHELY 1909)
Darevskia bithynica tristis (LANTZ & CYRÉN 1936) 
Common NamesG: Grusinische Eidechse, Kielschwanz-Felseidechse 
SynonymLacerta saxicola var. bithynica MÉHELY 1909: 495, 537
Lacerta saxicola bithynica — LANTZ & CYRÉN 1936: 178
Lacerta (Darevskia) rudis bithynica — SINDACO & JEREMČENKO 2008
Darevskia bithynica bithynica — ARRIBAS et al. 2013
Darevskia bithynica bithynica — ARRIBAS et al. 2022

Darevskia bithynica tristis (LANTZ & CYRÉN, 1936)
Lacerta saxicola tristis LANTZ & CYRÉN, 1936: 176
[Darevskia] saxicola tristis — DAREVSKY 1967t
Darevskia rudis tristis — BÖHME & BISCHOFF 1984
Lacerta (Darevskia) rudis tristis — SINDACO & JEREMČENKO 2008
Darevskia bithynica tristis — ARRIBAS et al. 2013
Darevskia bithynica tristis — ARRIBAS et al. 2022 
DistributionC/E Great Caucasus

Type locality: “Bithynischer Olymp bei Brussa”; “Amasia (Klein-Asien)”; lectotype locality: Turkey, Bursa Province, Mountain Bithynian Olympus (= Uludağ) in vicinity of Bursa (= Prusa), 1600 m asl, 40°09 ́N 29°07 ́E.

tristis: N Turkey; lectotype locality: Turkey, Sakarya Provinz, Ravine Lafaka Dere (= Lafakr-Dere, Maden Deresi?), fide Doronin 2017  
Reproduction 
TypesLectotype: ZISP 17128, designated by Doronin 2017.
Holotype: MZUT (Museo ed Istituto di Zoologia Sistematica, Torino) 2737:1
Lectotype: GNM 2481 (Göteborg Museum), designated by Darevsky 1967 [tristis] 
Diagnosis 
CommentSynonymy partly after KHALIKOV & ANANJEVA (pers. comm.). RYABININA et al. (2002) question the separation of bischoffi and obscura.

NCBI taxon ID: 1907507 [bithynica]
NCBI taxon ID: 1612236 [tristis] 
EtymologyNamed after the type locality. 
References
  • Arnold, E.N.; Arribas, O. & Carranza, S. 2007. Systematics of the Palaearctic and Oriental lizard tribe Lacertini (Squamata: Lacertidae: Lacertinae), with descriptions of eight new genera. Zootaxa 1430: 1-86 - get paper here
  • Arribas, O. J. 1999. Phylogeny and relationships of the mountain lizards of Europe and Near East (Archaeolacerta MERTENS, 1921, sensu lato) and their relationships among the eurasian lacertid radiation. Russ. J. Herpetol. 6 (1): 1-22 - get paper here
  • ARRIBAS, O., CANDAN, K., KORNILIOS, P., AYAZ, D., KUMLUTAŞ, Y., GÜL, S., ... & ILGAZ, Ç. 2022. Revising the taxonomy of Darevskia valentini (Boettger, 1892) and Darevskia rudis (Bedriaga, 1886)(Squamata, Lacertidae): a Morpho-Phylogenetic integrated study in a complex Anatolian scenario. Zootaxa 5224 (1): 1-68 - get paper here
  • ARRIBAS, OSCAR; ÇETİN ILGAZ, YUSUF KUMLUTAŞ, SALİH HAKAN DURMUŞ, AZİZ AVCI & NAZAN ÜZÜM 2013. External morphology and osteology of Darevskia rudis (Bedriaga, 1886), with a taxonomic revision of the Pontic and Small-Caucasus populations (Squamata: Lacertidae). Zootaxa 3626 (4): 401–428 - get paper here
  • Bedriaga, J. von 1886. Beiträge zur Kenntnis der Lacertiden-Familie (Lacerta, Algiroides, Tropidosaura, Zerzumia, Bettaia). Abh. senckenb. naturf. Ges. (Frankfurt) 14: 17-444 - get paper here
  • Bird, C. G. 1936. The distribution of reptiles and amphibians in Asiatic Turkey with notes on a collection from the Vilayets of Adana, Gaziantep and Malatya. Ann. Mag. Nat. Hist. (10) 18: 257-281. - get paper here
  • Bischoff, W. 1974. Echsen des Kaukasus, Teil 6. Die Kielschwanz-Felseidechse Lacerta rudis Bedriaga 1886. Aquar. Terr. Bl. 21, 274-278.
  • Bischoff, W. 2002. Auf der Suche nach Darevskia valentini (BOETTGER 1892) in Georgien - Impressionen aus einem wenig bekannten Land. Elaphe 10 (3): 49-61
  • Böhme,W. & Bischoff, W. 1984. Lacerta rudis - Kielschwanz-Felseidechse. In: Böhme,W. (ed.), Handbuch der Reptilien und Amphibien Europas, Band 2/I., Echsen II (Lacerta). Aula-Verlag Wiesbaden, pp. 332-344
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  • Budak,Abidin & Böhme,Wolfgang 1978. Über die rudis-Gruppe des Lacerta saxicola-Komplexes in der Türkei, I (Reptilia: Sauria: Lacertidae). Annalen des Naturhistorischen Museums in Wien 81 [1977]: 273-281 - get paper here
  • Darevsky, I. S. and Eiselt, J. 1980. Neue Felseneidechsen (Reptilia: Lacertidae) aus dem Kaukasus und aus der Turkei. Amphibia-Reptilia 1: 29-40. - get paper here
  • Darevsky, I.S. & Tuniyev,B.S. 1997. A new species from Lacerta saxicola group - Lacerta dryada sp. nov. (Sauria: Lacertidae) and some comments relative to Lacerta clarkorum Darevsky & Vedmederja 1977. Russ. J. Herpetol. 4 (1): 1-7 - get paper here
  • Darevsky,I.S. 1967. Rock lizards of the Caucasus: Systematics, ecology and phylogenesis of the polymorphic groups of Caucasian rock lizards of the the subgenus Archaeolacerta (Skal'nye Yashcheritsy Kavkasa; Izdat elst'vo Nauka, Leningrad). Leningrad, Nauka (English translation 1978): Ind. Nat. Sci. Doc. Centre; New Delhi, 276 pp. - get paper here
  • Doronin, I.V. 2017. REVIEW OF TYPE SPECIMENS OF THE ROCK LIZARDS OF DAREVSKIA (RUDIS) COMPLEX (SAURIA: LACERTIDAE) [in Russian]. Труды Зоологического института РАН 321 (3): 339–361 - get paper here
  • Eiselt,J. & Darevsky,I. 1991. Lacerta rudis chechenica ssp.n. aus dem Kaukasus (Reptilia: Lacertidae). Ann. Naturhist. Mus. Wien 92: 15-29 - get paper here
  • Engelmann, W.E. et al. 1993. Lurche und Kriechtiere Europas. Neumann Verlag (Radebeul, Germany), 440 pp.
  • FRANZEN, M. 1991. Beobachtungen zur Phytophagen Ernährung von Lacerta rudis und Lacerta clarkorum. Die Eidechse 2 (2): 22-23 - get paper here
  • Fu, JINZHONG AND ROBERT W. MURPHY 1997. Toward the phylogeny of caucasian rock lizards: implications from mitochondrial DNA gene sequences (Reptilia: Lacertidae). Zoological Journal of the Linnean Society (1997), 121: 463–477. - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Grechko VV, Ciobanu DG, Darevsky IS, Kosushkin SA, Kramerov DA. 2006. Molecular evolution of satellite DNA repeats and speciation of lizards of the genus Darevskia (Sauria: Lacertidae). Genome 49(10):1297-307. - get paper here
  • Grechko, V.V.; A. A. Bannikova, S. A. Kosushkin, N. L. Ryabinina, K. D. Milto, I. S. Darevsky and D. A. Kramerov 2007. Molecular Genetic Diversification of the Lizard Complex Darevskia raddei (Sauria: Lacertidae): Early Stages of Speciation. Molecular Biology, 41 (5): 764–775 - get paper here
  • Koç, Halime; Bilal Kutrup, Ufuk Bülbül, Muammer Kurnaz 2021. The Allelic Variants in Microsatellite Loci and Phylogenetic Relationships of Darevskia rudis (Bedriaga, 1886) and D. bithynica (Méhely, 1909) Based on Mitochondrial DNA in Turkey. Russian Journal of Herpetology 28 (2): 73-88 - get paper here
  • Langerwerf, B. 1978. Kaukasusrotshagedissen als ideale terrariumdieren. Lacerta 36 (12): 196-204 - get paper here
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  • MacCulloch, Ross D.;Fu, Jinzhong;Darevsky, Ilya S.;Danielyan, F.D.;Murphy, Robert W. 1995. Allozyme variation in three closely related species of Caucasian rock lizards (Lacerta). Amphibia-Reptilia 16 (4): 331-340 - get paper here
  • MANTEL, P. 1994. Bemerkenswerte Temperaturen bei der Inkubation von Eiern von Lacerta rudis bischoffi. Die Eidechse 5 (13): 4-7 - get paper here
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  • Murphy, R.W.; Fu,J.; MacCulloch, R.D.; Darevsky,I.S. & Kupriyanova,L.A. 2000. A fine line between sex and unisexuality: the phylogenetic constraints on parthenogenesis in lacertid lizards. Zool. J. Linnean Soc. 130: 527-549 - get paper here
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  • Ryabinin, D. M.;Grechko, V. V.;Darevsky, I. S.; Ryskov, A. P.; Semenova, S. K. 1996. Comparative study of DNA repetitive sequences by means of restriction endonucleases among populations and subspecies of some lacertid lizard species. Russ. J. Herpetol. 3 (2): 178-185 - get paper here
  • Ryabinina, N. L., A. A. Bannikova, S. A. Kosushkin, D. G. Ciobanu, K. D. Milto, B. S. Tuniyev, V. F. Orlova, V. V. Grechko and Illya S. Darevsky. 2003. Estimation of the subspecific level of differentiation in Caucasian lizards of the genus Darevskia (Syn. "Lacerta saxicola complex," Lacertidae, Sauria) using genome DNA Markers. Russ. J. Herpetol. 9 (3): 185-194 [2002] - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
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