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Delma butleri STORR, 1987

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Higher TaxaPygopodidae, Gekkota, Sauria, Squamata (lizards) 
Subspecies 
Common NamesUnbanded Delma, Butler's Legless Lizard, Spinifex Snake-lizard 
SynonymDelma butleri STORR 1987: 346
Delma haroldi STORR 1987: 350
Delma butleri — KLUGE 1993
Delma butleri — COGGER 2000: 286
Delma haroldi — MARYAN et al. 2007
Delma haroldi — WILSON & SWAN 2010: 130
Delma butleri — WILSON & SWAN 2010 
DistributionAustralia (New South Wales, Northern Territory, S Queensland, South Australia, Victoria, Western Australia)

Type locality: 5 km SE Gnaraloo, WA, in 23°52'S 113°32'E.

haroldi: Australia (SW Northern Territory, Western Australia);Type locality: 1 km NNW Daroo, WA, in 22°41'S, 115°20'E.  
Reproductionoviparous 
TypesHolotype: WAM R76742
Holotype: WAM R85094 [haroldi] 
DiagnosisDiagnosis: A small Delma without dark or pale bands across top of head and neck. Distinguishable from D. nasuta by fewer and differently patterned loreals, shorter and darker snout, dark brown upper labials variably marked with white (rather than brownish white, narrowly edged above with dark brown), dorsals dark brown finely edged with blackish (rather than pale brown, spotted dark brown) and ventrals and subcaudals never dark-edged. Distinguishable from D. grayii G.M. Storr by finely dark-edged dorsals and little or no development of ventrolateral vertical barring behind ear.

Diagnosis (haroldi): A small Delma with two pairs of supranasals, fourth upper labial below eye, usually 16 midbody scale rows, and with very narrow pale bands on head and neck but no dark bands. Most like faintly patterned examples of D. borea but distinguishable by pale bands on head and neck being wavy (rather than straight), by additional pale band(s) on side of head between postocular and auricular bands, and by dark spot at apex of anterior dorsals. 
CommentLimbless.

Synonymy: Shea (1991) considered D. haroldi a junior synonym of D. butleri. which was confirmed by Brennan et al. 2016), also it has been considered as valid by various recent authors. The range of D. haroldi lies within the range of D. butleri.

NCBI taxonID: 207581 [haroldi] 
EtymologyAfter naturalist W.H. (Harry) Butler, who was first to collect this species for the Western Australian Museum (on 30 December 1961 at Queen Victoria Spring).

Delma haroldi was named after naturalist Gregory Harold, co-collector of the holotype. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Brennan, Ian G.; Aaron M. Bauer, Todd R. Jackman 2015. Mitochondrial introgression via ancient hybridization, and systematics of the Australian endemic pygopodid gecko genus Delma. Molecular Phylogenetics and Evolution 94 (2016) 577–590
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp.
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Ellis, Ryan J.; Paul Doughty and Aaron M. Bauer 2018. An annotated type catalogue of the geckos and pygopods (Squamata: Gekkota: Carphodactylidae, Diplodactylidae, Gekkonidae, Pygopodidae) in the collection of the Western Australian Museum Records of the Western Australian Museum 33: 051–094 - get paper here
  • Jennings, W.B.; Pianka, E.R. & Donnellan, S. 2003. Systematics of the lizard family Pygopodidae with implications for the diversification of Australian temperate biotas. Systematic Biology 52: 757-780 - get paper here
  • Maryan, B. 2005. A HERPETOFAUNA HOTSPOT, THE CENTRAL WEST COAST OF WESTERN AUSTRALIA. Western Australian Naturalist 25 (1): 1-24
  • Maryan,B., Aplin,K., & Adams,M. 2007. Two new species of the Delma tincta group (Squamata: Pygopodidae) from northwestern Australia. Rec. West. Austr. Mus. 23: 273-305 - get paper here
  • Pianka, E. R. 1969. Habitat specificity, speciation, and species density in Australian desert lizards. Ecology 50 (3): 498-502 - get paper here
  • Pianka, E. R. 1986. Ecology and Natural History of Desert Lizards. Analyses of the Ecological Niche and Community Structure. Princeton University Press, Princeton, New Jersey
  • Pianka, E. R. 2010. Notes on the ecology of four species of pygopodid lizards in the Great Victoria Desert. Western Australian Naturalist 27: 175-181
  • Shea, G. M. 1991. Revisionary notes on the genus Delma (Squamata: Pygopodidae) in South Australia and the Northern Territory. Rec. South Austral. Mus. 25: 71-90 - get paper here
  • Storr, G. M. 1987. Three new legless lizards (Pygopodidae) from Western Australia. Rec. West. Austr. Mus. 13 (4): 345—355. - get paper here
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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