Dendrophidion crybelum CADLE, 2012

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Higher Taxa	Colubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies
Common Names
Synonym	Dendrophidion crybelum CADLE 2012 Dendrophidion vinitor — SAVAGE 1973: 14 (part.) Dendrophidion vinitor — SAVAGE 1980: 92 (part.) Dendrophidion vinitor — COTT et al., 1983: 372 (part.) Dendrophidion vinitor — SAVAGE & VILLA 1986: 17, 148, 169 (part.) Dendrophidion vinitor — LIEB 1988: 171 (part.) Dendrophidion vinitor — VILLA et al., 1988: 63 (part.) Dendrophidion vinitor — LIEB 1991: 522.1–522.2 (part.) Dendrophidion vinitor — AUTH 1994: 16 (part.) Dendrophidion vinitor — LEE 1996, fig. 164 (part.) Dendrophidion vinitor — SAVAGE 2002: 655–656 (part.) Dendrophidion vinitor — GOLDBERG 2003: 298– 300 (part.) Dendrophidion vinitor — STAFFORD 2003: 111 (part.) Dendrophidion vinitor — SOLÓRZANO 2004: 236–239 (part.) Dendrophidion vinitor — GUYER & DONNELLY 2005: 185 (part.) Dendrophidion vinitor — MCCRANIE et al., 2006: 147– 148 (part.) Dendrophidion vinitor — KÖHLER 2008: 215 (part.) Dendrophidion vinitor — SAVAGE & BOLAÑOS 2009: 14 (part.) Dendrophidion vinitor — MCCRANIE 2011: 111 (part). Dendrophidion crybelum — WALLACH et al. 2014: 225
Distribution	Costa Rica, Panama Type locality: Finca Las Cruces, near San Vito de Java, 4 km S San Vito, 1,200 m elevation, Puntarenas Province, Costa Rica
Reproduction	oviparous (not imputed, fide Zimin et al. 2022)
Types	Holotype: LACM 148599, an adult male. Collected in September 1972 by James E. DeWeese and Ron T. Harris. Figs. 2C, 12, 14B in Cadle 2012.
Diagnosis	Diagnosis: CADLE 2012: 210.
Comment	Distribution: not known from Panama with certainty, but the closest locality in Costa Rica is only ~1 km away from the Panamanian border, hence most likely in Panama too (S. Lotzkat, pers. comm., 23 Dec 2015). Habitat: partly arboreal (Harrington et al. 2018).
Etymology	The species name is derived from the Greek adjective krybelos meaning ‘‘hidden’’ or ‘‘secret.’’ Transliteration to Latin yields crybelum (with neuter gender ending to agree with the neuter generic name). It recognizes the cryptic nature of this species and the fact that it remained unrecognized for so long in a well-studied herpetofauna.
References	Cadle, John E. 2012. Cryptic Species Within the Dendrophidion vinitor Complex In Middle America (Serpentes: Colubridae). Bull. Mus. Comp. Zool. Harvard 160 (4): 183-240. - get paper here Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here Morato, Sérgio Augusto Abrahão; Guilherme Nunes Ferreira; Michela Rossane Cavilha Scupino (eds.) 2018. Herpetofauna da Amazônia Central: Estudos na FLONA de Saracá-Taquera. Curitiba, Pr: STCP Engenharia de Projetos Ltda.; Porto Trombetas, Pa: MRN – Mineração Rio do Norte S.A., 2018.<br />210p. - get paper here Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp. Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
External links	https://www.facebook.com/groups/stshc/posts/341914391366499 Google images

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