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Diploderma fasciatum (MERTENS, 1926)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Banded japalure
Chinese: 横纹龙蜥 
SynonymJapalura fasciata MERTENS 1926: 146
Japalura fasciata — SMITH 1935: 176
Japalura szechwanensis HU & ZHAO 1966 (fide OTA 2000, WANG et al. 2022)
Japalura szechwanensis — KÜHNEL et al. 1995
Japalura fasciata — BARTS & WILMS 2003
Japalura szechwanensis — LIVIGNI 2013: 414
Japalura szechwanensis — RAO et al. 2017
Japalura jinggangense WANG et al. 2017: 62 (fide WANG et al. 2022)
Diploderma fasciatum — WANG et al. 2018
Diploderma szechwanensis — GAO et al. 2022 
DistributionIndochina (Tonkin)

szechwanensis: China (Sichuan, Guizhou, Guangxi, Hunan, Yunnan). Type locality: Peng Hsien (= Peng Xien) and Tianquan Xien (= Tien Tsuan Hsien), Sichuan (= Szechwan), China.

Type locality: apparently not known. The type specimen was included in a collection purchased by the Senckenberg Museum from H. Fruhstorfer as of Tonkin (N Vietnam) without detailed type locality [BOETTGER 1904].  
Reproductionoviparous 
TypesHolotype: SMF 9871, male (formerly 4310,1a)
Holotype: CIB 613047, Paratype: CIB 625131, male, Tien Tsuan Hsien, 1,400 m asl. (= Tianquan Xien), Szechwan, China, Aug. 1962 [szechwanensis] 
DiagnosisDiagnosis: Diploderma fasciatum can be differentiated from congeners by a combination of the following characters: (1) body size medium to large, SVL up to 89.9 mm in adults; (2) tail length moderate, TAL 170.2–201.8% SVL in males, 172.2–196.6% in females; (3) forelimb length moderate, FLL 38.5–50.6% SVL; (4) hindlimb relatively short, HLL 55.9– 71.6% SVL; (5) nuchal crests much differentiated from dorsal, large, triangular in shape, CL 7.2–14.0% HL; (6) enlarged scales of dorsum highly differentiated from background scales, conical or sub-pyramidal, each bearing single, distinct medial keel; (7) post-temporal, post-tympanic, and post-rictal conical scales numerous and strongly developed; (8) tympa- num concealed; (9) ventral head scales heterogeneous in size, with enlarged gular scales scattered posteriorly and ventro- laterally; (10) F4S 18–25, T4S 21–29; (11) MD 34–49; (12) NSL mostly 0, rarely 1; (13) suborbital scale rows 3, middle row much enlarged; (14) gular pouch moderately developed in life, absent after preservation; (15) dorsolateral stripes absent in both sexes; (16) single hourglass-shaped, Light Greenish Yellow (Color 87) to Chartreuse (Color 89) transverse pattern present on dorsal midbody in both sexes, more profound in males; (17) distinct gular spots absent in both sexes; and (18) tongue and oral cavity Light Sulphur Yellow (Color 93) to Light Chrome Orange (Color 76), with dark gray speckles present towards the posterior lateral sides of oral cavity. (Wang et al. 2022)


Additional details (3102 characters) are available for collaborators and contributors. Please contact us for details. 
CommentSimilar species: D. grahami (which may be a synonym) 
References
  • ANANJEVA, Natalia B.; Xianguang GUO and Yuezhao WANG 2011. Taxonomic Diversity of Agamid Lizards (Reptilia, Sauria, Acrodonta, Agamidae) from China: A Comparative Analysis. Asian Herpetological Research 2 (3): 117-128 - get paper here
  • Barts, M. & Wilms, T. 2003. Die Agamen der Welt. Draco 4 (14): 4-23 - get paper here
  • Bobrov V.V., Semenov D.V. 2008. Lizards of Vietnam [in Russian]. Moscow, 236 pp.
  • Boettger, O. 1904. Museums-Bericht. I. Zoologische Sammlung. 4. Die Reptilien- und Batrachiersammlung. Ber. Senckenb. Naturforsch. Ges. Frankfurt am Main 1904: 126-131 - get paper here
  • DENZER, WOLFGANG; ULRICH MANTHEY, PATRICK D CAMPBELL 2019. Catalogue of type specimens of the agamid lizard genus Japalura s. l. (Squamata: Agamidae: Draconinae). Zootaxa 4612 (1): 109-125 - get paper here
  • Hu, Shu-QIN & Zhao, Ermi 1966. Three new species of reptiles from Szechwan (in Chinese with English abstract). Acta Zootaxon. Sinica, Peking, 3 (2): 158-164
  • LiVigni, F. (ed.) 2013. A Life for Reptiles and Amphibians, Volume 1. Chimaira, Frankfurt, 495 pp. - get paper here
  • Manthey U 2010. Agamid Lizards of Southern Asia. Draconinae 2 -Leiolepidinae. Edition Chimaira, Terralog 7b, Frankfurt, 168 pp.
  • MANTHEY, ULRICH; WOLFGANG DENZER, HOU MIAN & WANG XIAOHE 2012. Discovered in historical collections: Two new Japalura species (Squamata: Sauria: Agamidae) from Yulong Snow Mountains, Lijiang Prefecture, Yunnan, PR China. Zootaxa 3200: 27–48 - get paper here
  • Mertens,R. 1926. Herpetologische Mitteilungen. X. Eine neue Japalura-Art. Senckenbergiana 8 (3-4): 146-149
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Ota. H. 2000. Japalura szechwanensis, a junior synonym of J. fasciata. Journal of Herpetology 34 (4): 611-614 - get paper here
  • RAO, Dingqi; Jens V. VINDUM, Xiaohui MA, Mingxia FU and Jeffery A. WILKINSON 2017. A New Species of Japalura (Squamata, Agamidae) from the Nu River Valley in Southern Hengduan Mountains, Yunnan, China. Asian Herpetological Research 8 (2): 86-95, DOI:10.16373/j.cnki.ahr.160053 - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Wang K, Lyu ZT, Wang J, Qi S, Che J 2022. Updated Checklist and Zoogeographic Division of the Reptilian Fauna of Yunnan Province, China. Biodiversity Science 30 (4): 21326, 1–31 - get paper here
  • Wang, K., Qi, S., Wang, J., Köhler, G., Lu, C. Q., Lyu, Z. T., ... & Che, J. 2022. Revision of the Diploderma fasciatum (Mertens, 1926) Complex (Reptilia: Agamidae: Draconinae). Ichthyology & Herpetology, 110(3), 511-525 - get paper here
  • Wang, Kai; Jing Che, Simin Lin, V Deepak, Datta-Roy Aniruddha, Ke Jiang, Jieqiong Jin, Hongman Chen, Cameron D Siler; 2018. Multilocus phylogeny and revised classification for mountain dragons of the genus Japalura s.l. (Reptilia: Agamidae: Draconinae) from Asia. Zoological Journal of the Linnean Society, , zly034, - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
 
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