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Djokoiskandarus annulata (DE JONG, 1926)

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Higher TaxaHomalopsidae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Trans-Fly Mangrove Snake, Banded watersnake 
SynonymCantoria annulata DE JONG 1926: 304
Cantoria annulata — DAAN & HILLENIUS 1966: 137
Djokoiskandarus annulata — MURPHY 2011
Cantoria annulata — WALLACH et al. 2014: 146
Djokoiskandarus annulata — MURPHY & VORIS 2014: 14 
DistributionIndonesia (Irian Jaya), Papua New Guinea

Type locality: Prins Frederik Hendrik Island, West New Guinea (about 7°56'S and 138°21'E).  
Reproductionovovivparous 
TypesHolotype: ZMA.RENA 11065 
DiagnosisDiagnosis (genus): Dorsal scale rows 21 at mid body; internasal separates prefrontal and nasals scales; frontal’s contact with the internasal makes this species readily distinguishable from all other homalopsids, including Cantoria violacea which has 19 scale rows at mid body and the prefrontal scales in broad contact. A number of species of Enhydris have 21 scale rows at mid body, but all of them have the nasal scales in contact. D. annulatus also has keeled scales in the first three dorsal scale rows just anterior to the vent. A male is 578 mm in total length with a 108 mm tail; four females were 277–548 mm in total length. See Murphy (2007) for a more exhaustive description.


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CommentHabitat: marine (sea snake).

Diet: crustaceans

Type species: Cantoria annulata DE JONG 1926: 304 is the type species of the genus Djokoiskandarus MURPHY 2011. 
EtymologyNamed after the diminutive form (“annul-”) of Latin “anus” = ring.

The genus is named in honour of Indonesian herpetologist Djoko Iskandar. Gender of genus is masculine. 
References
  • Daan, S. & Hillenius,D. 1966. Catalogue of the type specimens of amphibians and reptiles in the Zoological Museum, Amsterdam. Beaufortia 13: 117-144
  • Heatwole, H. 1999. Sea snakes, 2nd ed. Krieger, Malabar, 148 pp. [review in Copeia 2000 (4): 1136] - get paper here
  • Jong,J.K. de 1926. Heurnia ventromaculata n.g. n. sp. und Cantoria annulata n. sp., wei neue Schlangen von Neu-Guinea. Zool. Anz. 67: 302-304
  • Murphy, J.C. 2011. THE NOMENCLATURE AND SYSTEMATICS OF SOME AUSTRALASIAN HOMALOPSID SNAKES (SQUAMATA: SERPENTES: HOMALOPSIDAE). Raffles Bulletin of Zoology 2011 59(2): 229–236 - get paper here
  • Murphy, J.C. & Voris, H.K. 2014. A Checklist and Key to the Homalopsid Snakes (Reptilia, Squamata, Serpentes), with the Description of New Genera. FIELDIANA: LIFE AND EARTH SCIENCES (8): 1–43 - get paper here
  • Murphy, John C. 2007. Homalopsid Snakes: Evolution in the Mud. Krieger Publishing, Malabar, Florida, 249 pp.
  • O’Shea,M. 1996. A Guide to the Snakes of Papua New Guinea. Independent Publishing, Port Moresby, xii + 239 pp. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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