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Draco cristatellus GÜNTHER, 1872

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Crested Flying Dragon
G: Leisten-Flugdrache 
SynonymDraco cristatellus GÜNTHER 1872
Draco cristatellus — BOULENGER 1885: 266
Draco cristatellus— DE ROOIJ 1915: 80
Draco cristatellus — INGER 1983
Draco cristatellus — MANTHEY & GROSSMANN 1997: 175
Draco cristatellus — MCGUIRE & KIEW 2001 
DistributionW Malaysia (Borneo: Sarawak)

Type locality: Borneo (Matang, Sarawak)  
Reproductionoviparous 
TypesHolotype: BMNH 1872.2.19.4 
DiagnosisDescription: Head large; snout as long as the diameter of the orbit; nostril lateral, directed outwards and upwards; tympanum partly scaly. Head-scales very small, keeled; a Y-shaped series of enlarged scales on the snout; a tubercle at the posterior corner of the orbit; eleven upper labials. Gular sac much longer than the head. A distinct nuchal fold. Dorsal scales smooth or faintly keeled, smaller than ventrals; no lateral series of large scales. Tail with a crest of long, separated, triangular scales, one time and a half as long as head and body. The fore limb extends beyond the tip of the snout, the hind limb reaches the axilla.

Coloration: Reddish-brown above, spotted with black; a black spot on the head between the orbits; membranes dark brown above, with irregular light longitudinal lines, below whitish with blackish spots along the margin. Gular sac yellow, the anterior edge brown. This is one of the few Draco species in which males and females have similar or identical patagial coloration (i.e. the color of the membranes that covers the elongated ribs).

Size: Length of head and body 78 mm; tail 112 mm (De Rooij 1915: 80). 
CommentSynonymy: Has been treated as a synonym of D. fimbriatus (HENNIG 1936, MUSTERS 1983, HONDA et al. 1999). Inger (1983) treated D. punctatus as a synonym of D. cristatellus. However, McGuire et al. 2001, 2018 recognized cristatellus as valid.

Distribution: not in Thailand (Chan-ard et al. 2015). Only known from a small area near Kuching. 
References
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Günther, A. 1872. On the reptiles and amphibians of Borneo. Proc. Zool. Soc. London 1872: 586-600. - get paper here
  • Inger,R.F. 1983. Morphological and ecological variation in the flying lizards (genus Draco). Fieldiana: Zoology, New Ser. 18: vi, 1-35 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • McGuire JA, Cotoras DD, O'Connell B, Lawalata SZS, Wang-Claypool CY, Stubbs A, Huang X, Wogan GOU, Hykin SM, Reilly SB, Bi K, Riyanto A, Arida E, Smith LL, Milne H, Streicher JW, Iskandar DT. 2018. Squeezing water from a stone: high-throughput sequencing from a 145-year old holotype resolves (barely) a cryptic species problem in flying lizards. PeerJ 6:e4470; DOI 10.7717/peerj.4470 - get paper here
  • McGuire, Jimmy A. & Heang, Kiew Bong 2001. Phylogenetic systematics of Southeast Asian flying lizards (Iguania: Agamidae: Draco) as inferred from mitochondrial DNA sequence data. Biological Journal of the Linnean Society 72: 203–229 - get paper here
  • McGuire, Jimmy A. and Robert Dudley 2011. The Biology of Gliding in Flying Lizards (Genus Draco) and their Fossil and Extant Analogs. Integrative and Comparative Biology, 51 (6): 983–990 - get paper here
 
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