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Draco maximus BOULENGER, 1893

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Common NamesE: Great Flying Dragon, Giant Gliding Lizard
G: Grosser Flugdrache 
SynonymDraco maximus BOULENGER 1893: 522
Draco intermedius WERNER 1910: 14 (fide HENNIG 1936)
Draco maximus — WERNER 1910: 16
Draco cryptotis DESPAX 1912
Draco maximus — DE ROOIJ 1915: 81
Draco cryptotis — DE ROOIJ 1915: 82
Draco intermedius — DE ROOIJ 1915: 86
Draco maximus — HENNIG 1936: 188
Draco maximus cryptotis — HENNIG 1936: 190
Draco maximus cryptotis — WERMUTH 1967: 52
Draco maximus maximus — GRANDISON 1972: 77
Draco maximus — MUSTERS 1983
Draco maximus — MANTHEY & GROSSMANN 1997: 170
Draco maximus — MCGUIRE & KIEW 2001
Draco maximus — TEYNIÉ et al. 2010
Draco maximus — GRISMER & QUAH 2019 
DistributionW Malaysia, Thailand, Indonesia (Sunda I, Sumatra, Natuna Island, Borneo)

cryptotis: Java; Type locality: “Archipel Malais”, restricted to Java by HENNIg 1936.

Type locality: Mount Dulit, Borneo (elevation 2000 f.h.)  
TypesType: BMNH 1946.8.13.41 (and possibly additional specimens). 
DiagnosisDiagnosis: Large species (maximum snout-vent length 145 mm); nostril directed upward; a more or less distinct series of scales forming alA-shaped figure on the snout; usually two median teeth in upper jaw; supralabials 13-16, smooth; no thornlike scale on supraciliary edge; tympanum partly covered with small scales or with small scales only; gular pouch without and lateral pouches with slightly enlarged scales; no nuchal crest; males with a nuchal fold; dorsals 226-269, subequal, smooth; six ribs in patagium; hindlimbs approximately as long as the distance between the limbs; no caudal crest (Musters 1983: 58). 
CommentThis is one of the few Draco species in which males and females have similar or identical patagial coloration (i.e. the color of the membranes that covers the elongated ribs). 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G.A. 1893. Description of new reptiles and batrachians obtained in Borneo by Mr. C. Hose and Mr. A. Everett. Proc. Zool. Soc. London 1893: 522-528 - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Denzer, W. & MANTHEY, U. 1991. A nominal checklist of the lizards inhabiting Peninsular Malaysia and Singapore. Raffles Bull. Zool., 39(2): 309-322 - get paper here
  • Despax, R. 1912. Sur trois collections de reptiles et de batraciens provenant de l’Archipel Malais. Bull. Mus. natl. Hist. nat. Paris 18: 198-205 - get paper here
  • Goldberg, Stephen R. and L. Lee Grismer. 2015. Notes on reproduction of seven species of Draco: D. abbreviatus, D. blanfordii, D. formosus, D. maculatus, D. maximus, D. sumatranus, D. taeniopterus (Squamata: Agamidae) from South-east Asia. Hamadryad 37 (1-2): 117-121 - get paper here
  • Grandison, A.G.C. 1972. The Gunong Benom Expedition 1967. 5. Reptiles and amphibians of Gunong Benom with a description of a new species of Macrocalamus. Bull. Br. Mus. nat. Hist. (Zool.), London, 23: 45-101. - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Hennig, W. 1936. Revision der Gattung Draco (Agamidae). Temminckia, Leiden 1: 153-220
  • Herlambang, A. E., Riyanto, A., Munir, M., Hamidy, A., Kimura, K., Eto, K., & Mumpuni, M. 2022. AFTER 16 YEARS: AN UPDATE CHECKLIST OF HERPETOFAUNA ON THE NATUNA ISLANDS, INDONESIA. TREUBIA, 49(2), 67-84 - get paper here
  • Inger,R.F. 1983. Morphological and ecological variation in the flying lizards (genus Draco). Fieldiana: Zoology, New Ser. 18: vi, 1-35 - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • McGuire, Jimmy A. & Heang, Kiew Bong 2001. Phylogenetic systematics of Southeast Asian flying lizards (Iguania: Agamidae: Draco) as inferred from mitochondrial DNA sequence data. Biological Journal of the Linnean Society 72: 203–229 - get paper here
  • McGuire, Jimmy A. and Robert Dudley 2011. The Biology of Gliding in Flying Lizards (Genus Draco) and their Fossil and Extant Analogs. Integrative and Comparative Biology, 51 (6): 983–990 - get paper here
  • Srichairat, Nattawut; Pattanee Jantrarotai, Prateep Duengkaeb, Yodchaiy Chuaynkern 2017. Identification key to species of the flying lizard genus Draco Linnaeus, 1758 (Squamata: Agamidae) in Thailand. Agriculture and Natural Resources - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Werner,F. 1910. Über neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. ii. Eidechsen. Jahrb. Hamburg. Wiss. Anst., vol. 27 (1909), suppl. no. 2, 1910, pp. 1-46; reprinted: 1910, Mitteil. Naturhist. Mus. Hamburg, vol. 27: 205-) - get paper here
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