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Draco quinquefasciatus HARDWICKE & GRAY, 1827

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Five-lined Flying Dragon, Five-banded Gliding Lizard
G: Fünfband-Flugdrache 
SynonymDraco 5 fasciata HARDWICKE & GRAY 1827: 219
Draco quinquefaseiata — HARDWICKE & GRAY 1835 (in error)
Draco quinquefasciatus — DUMÉRIL & BIBRON 1837: 455
Draco quinquefasciatus — BOULENGER 1885: 269
Draco quinquefasciatus — DE ROOIJ 1915: 81
Draco quinquefasciatus — TAYLOR 1963: 844
Draco quinquefasciatus longibarba HENNIG 1936
Draco quinquefasciatus longibarba — WERMUTH 1967: 53
Draco quinquefasciatus — GRANDISON 1972: 78
Draco quinquefasciatus — MUSTERS 1983
Draco quinquefasciatus longibarba — MANTHEY 1983
Draco quinquefasciatus — MANTHEY & GROSSMANN 1997: 173
Draco quinquefasciatus — COX et al. 1998: 103
Draco quinquefasciatus — MANTHEY & SCHUSTER 1999: 56
Draco quinquefasciatus longibarba — MALKMUS et al. 2002: 239
Draco quinquefasciatus — GRISMER 2011
Draco quinquefasciatus — GRISMER & QUAH 2019 
DistributionS Thailand, West Malaysia, Singapore (HR 33: 65),
Indonesia (Sumatra, P. Singkep, P. Belitung, Borneo, Natuna islands)

longibarba: Brunei, Borneo; Type locality: Baram, Sarawak, Borneo.

Type locality: restricted to Penang by HENNIG 1936  
Reproductionoviparous 
TypesType: BMNH 1946.8.27.4 (and possibly additional specimens).
Holotype: ZRC = RMS [longibarba] 
DiagnosisDiagnosis: Large or moderately sized species (maximum snout-vent length 106 mm); nostril directed upward; no A-shaped figure on the snout; usually two median teeth in upper jaw; supralabials 11-15, keeled; no thornlike scale on supraciliary edge; tympanum covered with small scales; gular pouch in males with slightly enlarged scales; lateral pouches with enlarged scales; nuchal fold in males; dorsals 178-213, smooth, subequal; usually six ribs in patagium; hindlimbs shorter than the distance between the limbs; no caudal crest (Musters 1983: 70). 
CommentThe original type locality was “India”, in error.

This is one of only 2 Draco species in which females have more colorful patagial coloration than males (i.e. the color of the membranes that covers the elongated ribs). 
References
  • Baker, Nick 2016. Five species of gliding lizard Draco spp. at Gunung Pulai, Johor, Peninsular Malaysia. SEAVR 2016: 110-112
  • Bong Heang, Kiew 1987. An annotated checklist of the herpetofauna of Ulu Endau, Johore, Malaysia. Malayan Nature Journal 41 (2-3): 413-423.
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, G.A. 1891. On new or little known Indian and Malayan reptiles and batrachians. Ann. Mag. nat. Hist. (6) 8: 288-292 - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • DAS, I. & LIM, K.K.P. 2001. Catalogue of herpetological types in the collection of the Raffles Museum of Biodiversity Research, National University of Singapore. Raffles Bull. Zool. 49 (1): 7-11 - get paper here
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Dring J C M 1979. Amphibians and reptiles from northern Trengganu, Malaysia, with descriptions of two new geckos: Cnemaspis and Cyrtodactylus. Bulletin of the British Museum (NATURAL HISTORY) ZOOLOGY 34 (5): 181-241 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Fehlandt, Lars 2017. Borneo – auf der Suche nach der Regenbogenkröte. Terraria-Elaphe 2017 (6): 62-65 - get paper here
  • FIGUEROA, ALEX; MARTYN E.Y. LOW, KELVIN K.P. LIM 2023. Singapore’s herpetofauna: updated and annotated checklist, history, conservation, and distribution. Zootaxa 5287 (1): 001–378 - get paper here
  • Grandison, A.G.C. 1972. The Gunong Benom Expedition 1967. 5. Reptiles and amphibians of Gunong Benom with a description of a new species of Macrocalamus. Bull. Br. Mus. nat. Hist. (Zool.), London, 23: 45-101. - get paper here
  • Gray. J. E. 1834. Illustrations of Indian Zoology, chiefly selected from the collection of Major - General Hardwicke. Vol. 2. London (1833-1834): 263 pp., 95 plates - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Günther, A. 1864. The Reptiles of British India. London (Taylor & Francis), xxvii + 452 pp. - get paper here
  • Hardwicke, Thomas & Gray, J.E. 1827. A synopsis of the species of saurian reptiles, collected in India by Major-General Hardwicke. Zoological Journal, London 3: 213-229 - get paper here
  • Hennig, W. 1936. Revision der Gattung Draco (Agamidae). Temminckia, Leiden 1: 153-220
  • Herlambang, A. E., Riyanto, A., Munir, M., Hamidy, A., Kimura, K., Eto, K., & Mumpuni, M. 2022. AFTER 16 YEARS: AN UPDATE CHECKLIST OF HERPETOFAUNA ON THE NATUNA ISLANDS, INDONESIA. TREUBIA, 49(2), 67-84 - get paper here
  • Husson, S.J.; S.H. Limin, Adul, N.S. Boyd, J.J. Brousseau, S. Collier, S.M. Cheyne, L.J. D’Arcy, R.A. Dow, N.W. Dowds, M.L. Dragiewicz, D.A. Ehlers Smith, Iwan, Hendri, P.R. Houlihan, K.A. Jeffers, B.J.M. Jarrett, I.P. Kulu, H.C. Morrogh-Bernard, S.E 2018. Biodiversity of the Sebangau tropical peat swamp forest, Indonesian Borneo. Mires and Peat 22 (05): 1–50 - get paper here
  • Inger,R.F. 1983. Morphological and ecological variation in the flying lizards (genus Draco). Fieldiana: Zoology, New Ser. 18: vi, 1-35 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Manthey,U. 1983. Exkursion am Mt. Kinabalu (4101 m), Nordborneo, Teil 3: Checkliste der Herpetofauna oberhalb 600 m ü. NN. Herpetofauna 5 (23): 20-31 - get paper here
  • Manthey,U. & SCHUSTER,N. 1999. Agamen, 2. Aufl. Natur und Tier Verlag (Münster), 120 pp. - get paper here
  • McGuire, Jimmy A. & Heang, Kiew Bong 2001. Phylogenetic systematics of Southeast Asian flying lizards (Iguania: Agamidae: Draco) as inferred from mitochondrial DNA sequence data. Biological Journal of the Linnean Society 72: 203–229 - get paper here
  • McGuire, Jimmy A. and Robert Dudley 2011. The Biology of Gliding in Flying Lizards (Genus Draco) and their Fossil and Extant Analogs. Integrative and Comparative Biology, 51 (6): 983–990 - get paper here
  • Nampochai, W., Rongchapho, P., Makchai, S., & Cota, M. 2021. Distribution of the Flying Lizard Genus Draco Linnaeus, 1758 (Squamata: Agamidae) of the Reference Collection of the Natural History Museum, National Science Museum, Thailand. The Thailand Natural History Museum Journal 15(1), 65-72 - get paper here
  • Srichairat, Nattawut; Pattanee Jantrarotai, Prateep Duengkaeb, Yodchaiy Chuaynkern 2017. Identification key to species of the flying lizard genus Draco Linnaeus, 1758 (Squamata: Agamidae) in Thailand. Agriculture and Natural Resources - get paper here
  • Stoliczka,F. 1873. Notes on some species of Malayan Amphibia and reptilia. J. Asiat. Soc. Bengal 42: 111-126 - get paper here
  • Sumarli, Alex X.; L. Lee Grismer, Shahrul Anuar, Mohd Abdul Muin and Evan S.H. Quah 2015. First report on the amphibians and reptiles of a remote mountain, Gunung Tebu in northeastern Peninsular Malaysia. Check List 11 (4): 1679 - get paper here
  • Taylor, E.H. 1963. The lizards of Thailand. Univ. Kansas Sci. Bull. 44: 687-1077. - get paper here
  • Teo, R.C.H. & N.J. Thomas 2019. Updated Inventory of Amphibians, Reptiles and Mammals of the Bukit Timah Nature Reserve, Singapore. Gardens’ Bulletin Singapore 71(Suppl. 1):145-183; doi: 10.26492/gbs71(suppl.1).2019-06 - get paper here
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
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