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Drymobius margaritiferus (SCHLEGEL, 1837)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
SubspeciesDrymobius margaritiferus margaritiferus (SCHLEGEL 1837)
Drymobius margaritiferus fistulosus SMITH 1942
Drymobius margaritiferus maydis VILLA 1968
Drymobius margaritiferus occidentalis BOCOURT 1890 
Common NamesE: margaritiferus: (Northern) Speckled Racer
G: Perlnatter 
SynonymColuber chiametla SHAW 1802 (nomen nudum ? fide WALLACH pers. comm.)
Natrix chiametla — SPIX 1824: 14
Herpetodryas margaritiferus SCHLEGEL 1837: 151
Leptophis margaritiferus — DUMÉRIL, BIBRON & DUMÉRIL 1854: 539
Zamenis tricolor HALLOWELL 1855: 34
Dromicus margaritiferus — GARMAN 1884: 58
Drymobius margaritiferus — BOULENGER 1894: 17
Drymobius margaritiferus margaritiferus — SMITH 1942: 383
Drymobius margaritiferus - VILLA et al. 1988
Drymobius margaritiferus margaritiferus — CONANT & COLLINS 1991: 190
Drymobius margaritiferus — LINER 1994
Drymobius margaritiferus margaritiferus — TENNANT & BARTLETT 2000: 345
Drymobius margaritiferus margaritiferus — LINER 2007
Drymobius margaritiferus — WALLACH et al. 2014: 246

Drymobius margaritiferus fistulosus SMITH 1942
Drymobius margaritiferus fistulosus SMITH 1942: 383
Drymobius margaritiferus fistulosus — SCHMIDT 1947
Drymobius margaritiferus fistulosus — DAVIS & SMITH 1953
Drymobius margaritiferus fistulosus — WILSON 1975
Drymobius margaritiferus fistulosus — LINER 2007

Drymobius margaritiferus maydis VILLA 1968
Drymobius margaritiferus maydis — WILSON 1975

Drymobius margaritiferus occidentalis BOCOURT 1890
Drymobius margaritiferus var. occidentalis BOCOURT 1890: 718
Drymobius margaritiferus occidentalis — SMITH 1942: 383
Drymobius margaritiferus occidentalis — MERTENS 1952: 64
Drymobius margaritiferus occidentalis — WILSON 1975
Drymobius margaritiferus occidentalis — LINER 2007 
DistributionUSA (S Texas) [Conant]
Mexico (incl. Tamaulipas, Querétaro, Puebla, Jalisco, Sinaloa, W Chihuahua, San Luis Potosí, Chiapas, Oaxaca, Hidalgo, Nuevo León), Panama, Guatemala, Honduras, Belize, El Salvador, Nicaragua, Costa Rica [Villa]
Colombia [Castro,F. (pers. comm.)]; elevation (Honduras): 0-1450 m

fistulosus: Mexico (incl. Michoacan, Nayarit, Sinaloa, Morelos); Type locality: Miramar,Nayarit, Mexico.

margaritiferus: S Texas; Type locality, "New Orleans" (in error); restricted to Veracruz (Mexico Smith, 1942); further restricted to Córdoba, Veracruz, Mexico (Smith and Taylor, 1950.

maydis: Nicaragua; Type Iocality, "Nicaragua: Departamento de Zelaya; Great Corn Island."

occidentalis: Guatemala, El Salvador; Type locality, "western [southern] versant of Guatemala near Volcán Atitlán.” Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
TypesHolotype unknown (margaritiferus)
Holotype: USNM 51480, female, collected by J. C. Thompson in 1913 [fistulosus]
Holotype: University of Costa Rica 999, aduIt male, collected by J. D. Villa, J. A. Roze, and R. G. Zweifel, January 1966 [maydis]
Syntypes: MNHN, Paris 7395 and 91·262, collector unknown, collected in 1866 [occidentalis] 
CommentRelative abundance in Honduras: rare

Type species: Herpetodryas margaritiferus SCHLEGEL 1837 is the type species of the genus Drymobius FITZINGER 1843. 
EtymologyThe generic name is derived from the Greek words drymos, meaning "oak wood or forest," and bios, meaning "life or manner of living," in reference to its habitat.
After the Latin “margarita” = pearl, and Latin “fer” = carrying. 
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