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Drymobius margaritiferus (SCHLEGEL, 1837)

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Higher TaxaColubridae, Colubrinae, Colubroidea, Serpentes (snakes) 
SubspeciesDrymobius margaritiferus margaritiferus (SCHLEGEL 1837)
Drymobius margaritiferus fistulosus SMITH 1942
Drymobius margaritiferus maydis VILLA 1968
Drymobius margaritiferus occidentalis BOCOURT 1890 
Common NamesE: margaritiferus: (Northern) Speckled Racer
G: Perlnatter 
SynonymColuber chiametla SHAW 1802 (nomen nudum ? fide WALLACH pers. comm.)
Natrix chiametla — SPIX 1824: 14
Herpetodryas margaritiferus SCHLEGEL 1837: 151
Leptophis margaritiferus — DUMÉRIL, BIBRON & DUMÉRIL 1854: 539
Zamenis tricolor HALLOWELL 1855: 34
Drymobius margaritiferus — BOULENGER 1894: 17
Drymobius margaritiferus - VILLA et al. 1988
Drymobius margaritiferus margaritiferus — CONANT & COLLINS 1991: 190
Drymobius margaritiferus — LINER 1994
Drymobius margaritiferus margaritiferus — TENNANT & BARTLETT 2000: 345
Drymobius margaritiferus margaritiferus — LINER 2007

Drymobius margaritiferus fistulosus SMITH 1942
Drymobius margaritiferus fistulosus — SCHMIDT 1947
Drymobius margaritiferus fistulosus — DAVIS & SMITH 1953
Drymobius margaritiferus fistulosus — WILSON 1975
Drymobius margaritiferus fistulosus — LINER 2007

Drymobius margaritiferus maydis VILLA 1968
Drymobius margaritiferus maydis — WILSON 1975

Drymobius margaritiferus occidentalis BOCOURT 1890
Drymobius margaritiferus var. occidentalis BOCOURT 1890: 718
Drymobius margaritiferus occidentalis — MERTENS 1952: 64
Drymobius margaritiferus occidentalis — WILSON 1975
Drymobius margaritiferus occidentalis — LINER 2007 
DistributionUSA (S Texas) [Conant]
Mexico (incl. Tamaulipas, Querétaro, Puebla, Jalisco, Sinaloa, W Chihuahua), Panama, Guatemala, Honduras, Belize, El Salvador, Nicaragua, Costa Rica [Villa]
Colombia [Castro,F. (pers. comm.)]; elevation (Honduras): 0-1450 m

fistulosus: Mexico (incl. Michoacan, Nayarit, Sinaloa, Morelos); Type locality: Miramar,Nayarit, Mexico.

margaritiferus: S Texas; Type locality, "New Orleans" (in error); restricted to Veracruz (Mexico Smith, 1942); further restricted to Córdoba, Veracruz, Mexico (Smith and Taylor, 1950.

maydis: Nicaragua; Type Iocality, "Nicaragua: Departamento de Zelaya; Great Corn Island."

occidentalis: Guatemala, El Salvador; Type locality, "western [southern] versant of Guatemala near Volcán Atitlán.” Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
TypesHolotype unknown [margaritiferus]
Holotype: USNM 51480, female, collected by J. C. Thompson in 1913 [fistulosus]
Holotype: University of Costa Rica 999, aduIt male, collected by J. D. Villa, J. A. Roze, and R. G. Zweifel, January 1966 [maydis]
Syntypes: MNHN, Paris 7395 and 91·262, collector unknown, collected in 1866 [occidentalis] 
CommentRelative abundance in Honduras: rare

Type species: Herpetodryas margaritiferus SCHLEGEL 1837 is the type species of the genus Drymobius FITZINGER 1843. 
  • Barbour,T. & Loveridge,A. 1929. Amphibians and reptiles [of the Corn Islands, Nicaragua]. Bull. Mus. comp. Zool. Harvard 69: 138-146 - get paper here
  • Boulenger, George A. 1894. Catalogue of the Snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Cobarrubias, Edgar Flores; Daniel Cruz-Sáenz, and David Lazcano 2012. Notes on the Herpetofauna of Western Mexico 6: Amphibians and Reptiles of Hostotipaquillo, Jalisco, Mexico. Bull. Chicago Herp. Soc. 47(2):21-26 - get paper here
  • Conant,R. & Collins,J.T. 1991. A Field Guide to Reptiles and Amphibians of Eastern/Central North America, 3rd ed. Houghton Mifflin (Boston/New York), xx + 450 p.
  • Crother, B. I. (ed.) 2012. Standard Common and Current Scientific Names for North American Amphibians, Turtles, Reptiles, and Crocodilians, Seventh Edition. Herpetological Circular 39: 1-92
  • Davis, William B.; Smith, Hobart M. 1953. Snakes of the Mexican state of Morelos Herpetologica 8: 133-149 - get paper here
  • Dixon, James R. 2000. Amphibians and reptiles of Texas, second edition. Texas A&M University Press, 421 pp.
  • Dixon, James R. and Julio A. Lemos-Espinal 2010. AMPHIBIANS AND REPTILES OF THE STATE OF QUERETARO, MEXICO. Tlalnepantla UNAM, 428 pp.
  • Guibé, J. & Roux-Estève, R. 1972. Les types de Schlegel (Ophidiens) présents dans les collections du Muséum National d'Histoire Naturelle de Paris. Zool. Meded. 47: 129-134 - get paper here
  • Hallowell, E. 1855. Contributions to South American Herpetology. Journal of the Academy of Natural Sciences, Philadephia (2) 3: 33-36 - get paper here
  • Heiko Werning 2013. Im Schlangenland – Ein Besuch bei Yvonne und Thomas Klesius. Terraria Elaphe 2013 (2): 88-93
  • Lee, J. C. 2000. A field guide to the amphibians and reptiles of the Maya world. Cornell University Press, Ithaca,
  • Lee, J.C. 1996. The amphibians and reptiles of the Yucatán Peninsula. Comstock, Cornell University Press, Ithaca, 500 pp.
  • Liner, Ernest A. 2007. A CHECKLIST OF THE AMPHIBIANS AND REPTILES OF MEXICO. Louisiana State University Occasional Papers of the Museum of Natural Science 80: 1-60
  • Martin, Plul S. 1958. A biogeography of reptiles and amphibians in the Gomez Farias Region, Tamaulipas, Mexico Miscellaneous publications, Museum of Zoology, University of Michigan (101): 1-102 + 7 plates - get paper here
  • McCranie J R 2011. The snakes of Honduras. SSAR, Salt Lake City, 725 pp.
  • McCranie, J. & Castañeda, F.E. 2005. The herpetofauna of Parque Nacional Pico Bonito, Honduras. Phyllomedusa 4 (1): 3-16 - get paper here
  • Mertens, R. 1952. Die Amphibien und Reptilien von El Salvador. Abh. senckenb. naturf. Ges. (Frankfurt) (No. 487): 120 pp.
  • Pérez-Santos,C. & Moreno,A.G. 1988. Ofidios de Colombia. Museo reegionale di Scienze Naturali, Torino, Monographie VI, 517 pp.
  • Porras, L.W. 2006. Die Schlangen des costa-ricanischen Trockenwaldes. Reptilia (Münster) 11 (61): 32-37 - get paper here
  • Savage, J.M. 2002. The Amphibians and Reptiles of Costa Rica: A Herpetofauna Between Two Continents, Between Two Seas. University of Chicago Press, 934 pp. [review in Copeia 2003 (1): 205]
  • Schlegel, H. 1837. Essai sur la physionomie des serpens. Partie Générale: xxviii +251 S. + Partie Descriptive: 606 S. + xvi. La Haye (J. Kips, J. HZ. et W. P. van Stockum) - get paper here
  • Schmidt, Karl P.;Shannon, Frederick A. 1947. Notes on amphibians and reptiles of Michoacan, Mexico. Zoological Series of Field Museum of Natural History 31 (9): 63-85
  • Smith, Hobart 1942. Mexican herpetological miscellany. Proc. US Natl. Mus. 92 (3153): 349-395 - get paper here
  • Smith, Hobart M. 1944. Snakes of the Hoogstraal Expeditions to northern Mexico. Zoological Series of Field Museum of Natural History 29 (8): 135-152 - get paper here
  • Spix, J.B. von. 1824. Serpentum brasiliensium species novae ou histoire naturelle des espèces nouvelles de serpens, recueillies et observées pendant le voyage dans l'intérieur du Brésil dans les années 1817, 1818, 1819, 1820, exécuté par ordre de sa majesté le Roi de Bavi Typis Franc. Seraph. Hübschmanni, Monachii viii + 75 pp.
  • Stejneger, L. 1891. Notes on some North American snakes. Proc. US Natl. Mus. 14 (876): 501-505 - get paper here
  • Stuart, L. C. 1935. A contribution to a knowledge of the herpetology of a portion of the savanna region of Central Peten, Guatemala University of Michigan Museum of Zoology Miscellaneous Publications 29: 1-56 - get paper here
  • Taylor,E.H. 1949. A preliminary account of the herpetology of the state of San Luis Potosi, Mexico. Univ. Kansas Sci. Bull. 33 (2): 169-215 - get paper here
  • Tennant, A. & Bartlett, R.D. 2000. Snakes of North America - Eastern and Central Regions. Gulf Publishing, Houston, TX, 588 pp.
  • Villa, J.D. 1968. A new colubrid snake from the Corn Islands, Nicaragua. Revista de Biologia Tropical 15 (1): 117-121 - get paper here
  • Webb, R.G. 1984. Herpetogeography in the Mazatlán-Durango Region of the Sierra Madre Occidental, Mexico. Vetrebrate Ecology and Systematics - A ribute to Henry S. Fitch; Museum of Natural History, University of Kansas, Lawrence, pp. 217-241
  • Werler, John E.&; James R. Dixon 2000. Texas Snakes. University of Texas Press, 544 pages
  • Wilson, L.D. & McCranie, J.R. 2003. The herpetofauna of the cloud forests of Honduras. Amphibian and Reptile Conservation 3 (1): 34-48 - get paper here
  • Wilson, Larry David 1975. Drymobius. Catalogue of American Amphibians and Reptiles (170): 1-2 - get paper here
  • Wilson,L.D. 1975. Drymobius margartitferus. Catalogue of American Amphibians and Reptiles No. 172: 1-2 - get paper here
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