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Dryophylax ramonriveroi (MANZANILLA & SÁNCHEZ, 2005)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Guianan Coastal House Snake 
SynonymThamnodynastes ramonriveroi MANZANILLA & SÁNCHEZ 2005
Thamnodynastes ramonriveroi — BAILEY & THOMAS 2007
Thamnodynastes nettingi (fide BAILEY & THOMAS 2007)
Thamnodynastes ramonriveroi — WALLACH et al. 2014: 718
Thamnodynastes ramonriveroi — NOGUEIRA et al. 2019
Thamnodynastes ramonriveroi — TREVINE et al. 2021
Dryophylax ramonriveroi — TREVINE et al. 2022 
DistributionVenezuela (Turimiquire massiff, elevation 0-1750 m), Suriname (Nieuw Nickerie and Paramaribo), Guyana (Demerara [Georgetown]), Brazil

Type locality: Cerro La Laguna, Turimiquire massif, Fre- ites, Anzoátegui, Venezuela.  
TypesHolotype: EBRG 3957, a 469 mm male 
DiagnosisDiagnosis: A northern dwarfed relative of Thamnodynastes strigatus, from which it differs chiefly in its smaller size, lesser development of para-cloacal tubercles in males, a higher modal maxillary tooth count, and in possessing enlarged basal hooks on the hemipenis.

Character table: Franco et al. 2017, Trevine et al. 2021: 238 (Table 1) 
CommentSympatric with T. pallidus. 
EtymologyNamed after Ramón Rivero, who for more than 20 years has enriched and maintained the reptile collection of the Rancho Grande Biological Station, making and allowing great contributions to the knowledge of the Venezuelan herpetofauna. 
  • Bailey, J.R.; R.A. Thomas 2007. A revision of the South American snake genus Thamnodynastes Wagler, 1830 (Serpentes, Colubridae, Tachymenini). II. Three new species from northern South America, with further descriptions of Thamnodynastes gambotensis Pérez-Santos and Moreno and Tha Memoria de la Fundación La Salle de Ciencias Naturales 166: 7-27 [2006]
  • Cole, Charles J.; Carol R. Townsend, Robert P. Reynolds, Ross D. MacCulloch, and Amy Lathrop 2013. Amphibians and reptiles of Guyana, South America: illustrated keys, annotated species accounts, and a biogeographic synopsis. Proceedings of the Biological Society of Washington 125 (4): 317-578; plates: 580-620 - get paper here
  • Hedges SB, Powell R, Henderson RW, Hanson S, and Murphy JC 2019. Definition of the Caribbean Islands biogeographic region, with checklist and recommendations for standardized common names of amphibians and reptiles. Caribbean Herpetology 67: 1–53
  • Manzanilla, J. & Sánchez 2005. Una nueva especie de Thamnodynastes (Serpentes: Colubridae) del macizo del Turimiquire, noreste de Venezuela. Memorias de la Fundación La Salle de Ciencias Naturales 161-162: 61-75 [2004]
  • Natera-Mumaw, Marco; Luis Felipe Esqueda-González & Manuel Castelaín-Fernández 2015. Atlas Serpientes de Venezuela. Santiago de Chile, Dimacofi Negocios Avanzados S.A., 456 pp. - get paper here
  • Nogueira, Cristiano C.; Antonio J.S. Argôlo, Vanesa Arzamendia, Josué A. Azevedo, Fausto E. Barbo, Renato S. Bérnils, Bruna E. Bolochio, Marcio Borges-Martins, Marcela Brasil-Godinho, Henrique Braz, Marcus A. Buononato, Diego F. Cisneros-Heredia, 2019. Atlas of Brazilian snakes: verified point-locality maps to mitigate the Wallacean shortfall in a megadiverse snake fauna. South American J. Herp. 14 (Special Issue 1):1-274 - get paper here
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Trevine, V. C., Grazziotin, F. G., Giraudo, A., Sallesbery‐Pinchera, N., Vianna, J. A., & Zaher, H. 2022. The systematics of Tachymenini (Serpentes, Dipsadidae): An updated classification based on molecular and morphological evidence. Zoologica Scripta - get paper here
  • TREVINE, VIVIAN C.; JOSÉ RANCÉS CAICEDO-PORTILLA, MARINUS HOOGMOED, ROBERT A. THOMAS, FRANCISCO L. FRANCO, GIOVANNA G. MONTINGELLI, MARIELA OSORNO-MUÑOZ, HUSSAM ZAHER 2021. A new species of Thamnodynastes Wagler, 1830 from western Amazonia, with notes on morphology for members of the Thamnodynastes pallidus group (Serpentes, Dipsadidae, Tachymenini). Zootaxa 4952 (2): 235–256 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
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