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Ebenavia inunguis BOETTGER, 1878

IUCN Red List - Ebenavia inunguis - Least Concern, LC

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Common NamesE: Madagascar Clawless Gecko
D: Pinselschwanzgecko 
SynonymEbenavia inunguis BOETTGER 1878
Ebenavia inunguis — BOULENGER 1885: 96
Ebenavia boettgeri BOULENGER 1885: 96
Ebenavia inunguis — KLUGE 1993
Ebenavia inunguis — GLAW & VENCES 1994: 263
Ebenavia inunguis — RÖSLER 1995: 114
Ebenavia inunguis — BROADLEY & HOWELL 1991: 9
Ebenavia inunguis — RÖSLER 2000: 78
Ebenavia inunguis — SPAWLS et al. 2018: 75 
DistributionNW Madagascar, Nossi Be

Type locality: Nossi Be = Nosy Bé  
TypesHolotype: SMF 8318, originally 4096a, adult male. Boettger (1878:10) explicitly stated that his description was based on a single individual. Boettger (1893:26) indicated that 4096a consisted of four individuals. Mertens (1967) selected the original holotype as the lectotype of a series that also comprised three adult females SMF 8332–8334, which were originally joined with SMF 8318 under the number 4096a. 
DiagnosisDiagnosis (genus). Leaf-toed (phyllodactyle) gekkonines with claws restricted to digits of pes of females; dorsolateral scales of head, body, and limbs multicarinate except few enlarged anterior plates on tip of snout of E. inunguis; variable number of multicarinate scales on tail and ventral surfaces. (Nussbaum & Raxworthy 1998) 
CommentSynonymy: Ebenavia boettgeri Boulenger, 1885 was considered a synonym of Ebenavia inunguis by Boulenger (1887) and subsequent authors but was resurrected as a valid species by Hawlitschek et al. 2018.

Type Species: Ebenavia inunguis BOETTGER 1878 is the type species of the genus Ebenavia BOETTGER 1878.

Taxonomy: Hawlitschek et al. 2016 provide evidence for the existence of a species complex.

Distribution: see map in Hawlitschek et al. 2018: 48 (Fig. 5). 
EtymologyThe specific name "inunguis", (Latin) for "without claws", is a misnomer as females have claws on their toes (but see description).

The genus Ebenavia was named after the collector Carl (sometimes written as Karl) Ebenau and is therefore formed neither from a Latin nor a Greek word. The fact that the name is consistently written as Ebenavia (as opposed to ‘Ebenauia’) throughout the paper suggests that the ‘u’ was intentionally changed to ‘v’. The reason for this change was not given by Boettger, but it may have been a latinisation of the name, or for greater euphony, or both. The gender of the genus was not mentioned by Boettger (1878) and can neither be derived from the species name inunguis nor from the later described taxa boettgeri (named after Oskar Boettger) and maintimainty (Malagasy adjective). Based on Article 30 (‘Gender of genus-group names’) of the International Code of Zoological Nomenclature (1999) and more specifically Article 30.2.4. (‘If no gender was specified or indicated, the name is to be treated as masculine, except that, if the name ends in -/a/ the gender is feminine, and if it ends in -/um/, -/on/, or -/u/ the gender is neuter.’). The gender of Ebenavia is feminine (from Hawlitschek et al. 2018). 
  • Andreone F., Randrianirina J., Jenkins P.D. & Aprea G. 2000. Species diversity of Amphibia, Reptilia and Lipotyphla (Mammalia) at Ambolokopatrika, a rainforest between the Anjanaharibe-Sud and Marojejy massifs, NE Madagascar. Biodiversity and Conservation 9: 1587–1622 - get paper here
  • Barbour, Thomas 1918. Vertebrata from Madagascar. 2. Amphibia and Reptilia. Bull. Mus. Comp. Zool. Harvard 61 (14): 479-489. - get paper here
  • Bauer A M. Good D A. Branch W R. 1997. The taxonomy of the Southern African leaf-toed geckos (Squamata: Gekkonidae), with a review of Old World "Phyllodactylus" and the description of five new genera. Proc. Cal. Acad. Sci. 49 (14): 447-497. - get paper here
  • Boettger, O. 1878. Die Reptilien und Amphibien von Madagascar. I. Studien über Reptilien aus Madagascar. II. Aufzählung der bis jetzt von Madagascar bekannt gewordenen Reptilien und Amphibien. III. Bemerkungen über die verwandtschaftlichen und geographischen Beziehunge Abh. senckenb. naturf. Ges. (Frankfurt) 11: 1-56 [1877] - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
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  • D’Cruze, N.; Köhler, J.; Franzen, M & Glaw, F. 2008. A conservation assessment of the amphibians and reptiles of the Forêt d’Ambre Special Reserve, north Madagascar. MADAGASCAR CONSERVATION & DEVELOPMENT 3 (1): 44-54 - get paper here
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  • Hawlitschek, O.; F. Glaw & D. Rödder 2012. Pemba – Herpetologische Fundgrube im Indischen Ozean. Reptilia (Münster) 17 (97): 97-109 - get paper here
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  • Zimmermann, H. 2001. Das erste “deutsche” Naturschutzgebiet in Madagaskar, der “Drachenbaum-Regenwald Maromiza bei Andasibe und seine Herpetofauna. Herpetofauna 23 (134): 28-34 - get paper here
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