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Echis romani TRAPE, 2018

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Higher TaxaViperidae, Viperinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymEchis romani TRAPE 2018 
DistributionSW Chad, S Niger, Nigeria, N Cameroon, NW Central African Republic

Type locality: Kumao, Chad, Lagone Oriental region, Baibokoum, 7°36’ N, 15° 36’ E)  
Reproduction 
TypesHolotype: MNHN-RA 2018.0006, male (formerly IRD 2013.N);
Paratypes: (n=24) MNHN 2018.0007, previously IRD 2026.N, MNHN 2018.0008, previously IRD 2157.N, MNHN 2018.0009, previously IRD 2135.N, MNHN 2018.0010, previously IRD 2038.N, MNHN 2018.0011, previously IRD 2100.N, MNHN 2018.0012, previously IRD 2230.N; IRD 2011.N, 2034.N, 2057.N, 2078.N, 2124.N, 2126.N, 2127.N, 2128.N, 2133.N, 2134.N, 2139.N, 2190.N, 2268.N, 2272.N, 2333.N and 2334.N, all collected by villagers from May 20 to 30, 2015 in the around Baïbokoum (07°35’-07°46’N / 15°35’-15°41’E) and IRD 2550.N and 2734.N collected by villagers in November 2015 in Laobida (09°12’N / 15°07’E) in Mayo Kebbi in Chad. 
DiagnosisDiagnosis: A large species (up to 715 mm in total length) genetically distinct from other species of the E. ocellatus complex and characterized by the combination of following characters: (1) a large number of ventral scales (148 – 151.9 – 156 in males and 152 – 159.9 – 168 in females), (2) simple subcaudal scales at the number from 25 to 29 in males (average 27.1) and from 18 to 23 in females (average 20.9), (3) 29 to 31 dorsal rows keeled in the middle of the body in males (average 30.7) and 31 to 33 in females (mean 31.3), (4) a large number of small scales surrounding each eye (mean 16.2), (5) dorsal coloration pattern with brown vertebral stripe dark regularly interspersed with light spots, on the sides dark spots with white ocelli most often located next to the medio-dorsal light spots and the presence three to six dark spots on each ventral scale. (Trape 2018) 
CommentDistribution: see map in Trape 2018: 31 (Fig. 15). 
EtymologyNamed after Benigno Roman for his contributions to African herpetology and ichthyology. 
References
  • Spawls, Stephen; Tomáš Mazuch & Abubakr Mohammad 2023. Handbook of Amphibians and Reptiles of North-east Africa. Bloomsbury, 640 pp. - get paper here
  • Trape, J.-F. 2018. Partition d’Echis ocellatus Stemmler, 1970 (Squamata, Viperidae), avec la description d’une espèce nouvelle. Bull. Soc. Herp. France 167: 13-34 - get paper here
  • Trape, J.-F. 2023. Guide des serpents d’Afrique occidentale, centrale et d’Afrique du Nord. IRD Éditions, Marseille, 896 pp.
  • Trape, Jean-François; Israël Demba Kodindo, Ali Sougoudi Djiddi, Joseph Mad-Toïngué & Clément Hinzoumbé Kerah 2020. The snakes of Chad: results of a field survey and annotated country-wide checklist. Bonn zoological Bulletin 69 (2): 367–393 - get paper here
 
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