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Emmochliophis fugleri FRITTS & SMITH, 1969

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Higher TaxaColubridae (Dipsadinae), Diaphorolepidini, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Pinchinda Snake 
SynonymEmmochliophis fugleri FRITTS & SMITH 1969
Emmochliophis fugleri — SHEIL 1998
Emmochliophis fugleri — WALLACH et al. 2014: 268 
DistributionW Ecuador

Type locality: Ecuador, Pichincha, 4 km E Rio Baba bridge, 24 km S Sto. Domingo de los Colorados, 600 m elevation.  
TypesHolotype: INHS (= UIMNH) 78795 
DiagnosisDescription (genus). Relatively small-sized (~250mm SVL) terrestrial snakes restricted to the Pacific Andean slopes of NW Ecuador, with a small number (<17) of maxillary teeth, 8 supralabials, 8 infralabials, fused prefrontals, internasals in contact, loreal absent, fewer than 150 ventrals, fewer than 100 subcaudals, dorsal scales in 19 rows without reduction, trunk vertebrae with lateral expansion of the zygapophyses, and expanded zygapophyses forming a rod-and-groove mechanism in Emmochliophis fugleri, but not in E. miops [PYRON et al. 2015: 125].

Description (species). A terrestrial snake from the Pacific Andean slopes of NW Ecuador, diagnosable by 16 maxillary teeth, 8 infralabials, 8 supralabials, 2 postoculars, internasals in contact, loreal absent, nuchal collar absent, 140 ventrals, 97 subcaudals, dorsal scales in 19 rows without reduction, strong keels, and zygapophyses expanded laterally forming rod–and–bar assembly. Type locality is surrounded by banana plantations. Little else is known about the habits or habitat of the species [PYRON et al. 2015: 125]. 
CommentKnown from only one (male) type specimen.

Type species: Emmochliophis fugleri FRITTS & SMITH 1969 is the type species of the genus Emmochliophis FRITTS & SMITH 1969. 
EtymologyFrom the Greek emmochlion for “a socket for a bar” and ophis for “snake,” referring to the unique interlocking vertebrae (Fritts and Smith 1969).
Species name after Dr. Charles Fugler, who collected the holotype. 
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Fritts, Thomas H. Smith, Hobart M. 1969. A new genus and species of snake from western Ecuador. Transactions of the Kansas Academy of Science 72 (1): 60-66 - get paper here
  • Maynard RJ, Culebras J, Kohn S, Guayasamin JM, Trageser SJ 2021. Finding a shadow in the dark: rediscovery of Fugler’s Shadow Snake (Emmochliophis fugleri Fritts & Smith, 1969) after 54 years, with comments on its conservation status, distribution, and the tribe Diaphorolepidini. Check List 17(1): 239-245 - get paper here
  • Pyron RA, Guayasamin JM, Peñafiel N, Bustamante L, Arteaga A 2015. Systematics of Nothopsini (Serpentes, Dipsadidae), with a new species of Synophis from the Pacific Andean slopes of southwestern Ecuador. ZooKeys 541: 109-147, doi: 10.3897/zookeys.541.6058 - get paper here
  • Sheil,C.A. 1998. Emmochliophis miops: redescription of Synophis miops (BOULENGER 1898). Journal of Herpetology 32 (4): 604-607 - get paper here
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
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