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Emydura subglobosa (KREFFT, 1876)

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Higher TaxaChelidae, Chelodininae, Pleurodira, Testudines (turtles) 
SubspeciesEmydura subglobosa subglobosa (KREFFT 1876)
Emydura subglobosa worrelli WELLS & WELLINGTON 1985
Emydura subglobosa angkibaanya JOSEPH-OUNI, MCCORD, CANN, SMALES & FREEMAN 2019 
Common NamesE: Red-bellied Short-necked Turtle
worrelli: Diamond-head Turtle
G: Rotbauch-Spitzkopfschildkröte 
SynonymEuchelymys subglobosa KREFFT 1876
Emydura albertisii BOULENGER 1888 (fide CANN 1978)
Emydura albertisii — PODLOUCKY 1984
Tropicochelymys goodei WELLS & WELLINGTON 1985 (fide IVERSON et al. 2001)
Emydura subglobosa — IVERSON 1986
Emydura subglobosa — COGGER 2000: 199
Emydura subglobosa subglobosa — GEORGES & THOMSON 2010

Emydura subglobosa worrelli WELLS & WELLINGTON 1985
Tropicochelymys worrelli WELLS & WELLINGTON 1985: 9
Tropicochelymys leichhardti WELLS & WELLINGTON 1985, nomen nudum
Emydura victoriae worrelli
Emydura worrelli — COGGER 2000: 740 (see comment)
Emydura subglobosa worrelli — GEORGES & ADAMS 1996
Emydura subglobosa worrelli — MCCORD et al. (2003)
Emydura worrelli — BONIN et al 2006
Emydura subglobosa worrelli — GEORGES & THOMSON 2010 
DistributionS Papua New Guinea,
Australia (Jardine River at the northern tip of the Cape York Penisula, Queensland)

Type locality: Naiabui, on Amama River, SE Papua New Guinea.

angkibaanya: Australia (N Queensland); 'Old River Crossing' , Jardine River, Cape York Peninsula, Queensland, Australia, 11°08'S, 142°21'E (fide Shaffer et al., 2009)  
Reproductionoviparous. 
TypesHolotype: MSNG 2320 (was CE; Museo Civico di Storia Naturale di Genova)
Holotype: AMS R53689 [worrelli]
Holotype. AMS R37669, adult female of 160 mm straight carapace length, collected 1972 by G. Webb and R. Shine. The prepared skull of this specimen is catalogued as AMS R37669.01. Paratypes: AMS R37666, juvenile; AMS R37668, adult male [angkibaanya] 
DiagnosisDescription and Diagnosis (angkibaanya). We follow the species description of Emydura subglobosa whose specific identity is revised and defined by Smales et al., 2019, characters of which therein documented (i.e. a species of the genus Emydura bearing the combination of light temporal/post-orbital stripe extending to tympanum; pale golden-yellow iris; anterior and posterior black spots form a horizontal bar through the iris; red coloration flush across the ventral carapace surfaces; oval to oblong shell with the carapace being dark-brown to black; etc.) extend to both the populations of E. subglobosa in southern New Guinea (the nominate form) and the Jardine River, Queensland, Australia (angkibaanya).
E. s. angkibaanya ssp. nov. is further diagnoseable by the following combination o f characters: a consistently oval-shaped carapace; an uniformly black or all dark brown carapace that can bear darker mottling and that lacks reddish to reddish-yellow lighter centers o f the carapacial scutes in life ; a throat pattern that is dark gray, often with a centralized reddish-yellow patch; a dark figure or wash over the bridges and ventral marginal scute surfaces that is retained in most sexually mature specimens; a dark blotch present at the center of the posterior mandibular skin; a bright yellow to greenish yellow iris, with some specimens bearing darker vermiculation; a yellowish-gray sclera often with darker markings; a yellow or orangish-yellow post-orbital stripe that consistently widens and makes contact to the antero-dorsal tympanum rim; and a horse-shoe shaped dark marking along the lower jawline/ throat skin border that continuously merges with the dark facial skin behind the lower jaw. All colors as in life (see Plates 4, 5 and 6; sketch of carapace shape presented in Figure 4). There are minimal morphological differences between the sexes, however we note that by personal observation, adult female specimens tend to retain the bold coloration of the ventral surfaces more vividly than male specimen do (field observations and plate 6) and that male specimens rarely exceed 150 mm straight carapace length as seen in Australian Museum paratype specimen R37668.

Comparisons (angkibaanya): Joseph-Ouni et al. 2019: 56. 
CommentSynonymy: Emydura albertisii has been included in E. kreffti (GOODE 1967) or in E. australis (PRITCHARD 1979). GOODE (1967) also synonymized E. schultzei with Emydura (= Elseya) novaeguineae and Emydura subglobosa and Emydura albertisii with Emydura krefftii. FRITZ & BAUR (1995) report a vital hybrid between Emydura subglobosa and Elseya novaeguineae.

Distribution: Map in MCCORD et al. (2003) and Joseph-Ouni et al. 2019 (Appendix, Fig. 5). WELLS & WELLINGTON (1985) separated the Batten Creek and MacArthur River, Northern Territory, populations as Tropicochelymys worrelli (fide KING & BURKE 1989).

Etymology: The subspecific name angkibaanya references the Australian Aboriginal dialectic word for 'rainbow' in the language of the Gudang people who are traditional inhabitants of the northern-most area of the Cape York Peninsula, in allusion to this taxon's bold facial and ventral coloration.

Habitat: freshwater (rivers, swamps) 
Etymology 
References
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