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Enyalioides binzayedi VENEGAS, TORRES-CARVAJAL, DURAN & DE QUEIROZ, 2013

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Higher TaxaHoplocercidae, Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common Names 
SynonymEnyalioides binzayedi VENEGAS, TORRES-CARVAJAL, DURAN & DE QUEIROZ 2013 
DistributionPeru (Picota)

Type locality: Chambirillo close to the Checkpoint 16 of the CAZNP (07°04’8.9”S, 76°00’51.2”W, 1122 m elevation), Provincia de Picota, Región San Martín, Perú Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproduction 
TypesHolotype: CORBIDI 08828 (Fig. 7), an adult male, collected on 30 October 2010 by P. J. Venegas.
Paratypes. CORBIDI 08827, an adult female collected on 2 November 2010 by P. J. Venegas; CORBIDI 08786, 08787, 08788, 08789, adult females collected on 21 January 2011 by P. J. Venegas and V. Duran; CORBIDI 09215, 09216, a juvenile male and adult female, respectively, collected on 6 May 2011 by P. J. Venegas and V. Duran. All paratypes are from the same locality as the holotype. 
CommentDiagnosis. Enyalioides binzayedi can be distinguished from other species of Enyal- ioides by the combination of the following characters: (1) scales posterior to the super- ciliaries forming a longitudinal row of strongly projecting scales across the lateral edge of the skull roof in adults of both sexes; (2) 31 or fewer longitudinal rows of strongly keeled dorsals in a transverse line between the dorsolateral crests at midbody; (3) ventral scales strongly keeled; (4) caudals increase in size posteriorly within each autotomic segment; (5) projecting scales on body or limbs absent; (6) vertebrals on neck more than five times the size of vertebrals between hind limbs in adult males.
A longitudinal row of strongly projecting scales along the lateral edge of the skull posterior and continuous with the superciliaries is also present in E. oshaughnessyi, which occurs west of the Andes in Ecuador and Colombia and differs from E. bin- zayedi in having smooth or slightly keeled dorsals. Species of Enyalioides occurring east of the Andes that share strongly keeled ventrals with E. binzayedi are E. azulae, E. cofanorum, E. microlepis, E. palpebralis, and E. rudolfarndti. All of these species either lack strongly projecting scales along the lateral edge of the skull roof (although they are slightly projecting in E. rudolfarndti) or have such scales but with a gap separating them from the superciliaries (E. palpebralis). Enyalioides azulae, E. cofanorum and E. microlepis differ further from E. binzayedi (character states in parentheses) in having more than 33 dorsal scales in a transverse line between the dorsolateral crests at mid- body (31 or fewer), a low vertebral crest (high, with vertebrals on neck more than four times the size of vertebrals between hind limbs in both sexes), and a black gular patch (absent). The new species can be also distinguished from E. palpebralis by lacking both a superciliary triangular flap that projects posterolaterally over each eye and a small gap in the vertebral crest in the neck region, and by having femoral pores. From E. rudol- farndti (character states in parentheses), E. binzayedi also differs in having a prominent medial keel on each dorsal scale (medial keel weak or absent), dorsals nearly homoge- neous in size (dorsals heterogeneous in size), and in lacking a round orange blotch in the antehumeral region (orange blotch present in adult males). 
EtymologyThe specific name is a noun in the genitive case and is a patronym honoring Sheikh Mohamed bin Zayed Al Nahyan, Crown Prince of Abu Dhabi and Deputy Supreme Commander of the UAE, who created the Mohamed bin Zayed Species Conservation Fund (MBZSCF) to support species conservation projects around the globe. Field surveys leading to the discovery of the two species reported on in this paper were supported by a grant from the MBZSCF. 
References
  • Venegas PJ, Torres-Carvajal O, Duran V, de Queiroz K 2013. Two sympatric new species of woodlizards (Hoplocercinae, Enyalioides) from Cordillera Azul National Park in northeastern Peru. ZooKeys 277: 69–90. doi: 10.3897/zookeys.277.3594 - get paper here
 
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