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Epictia vindumi WALLACH, 2016

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Higher TaxaLeptotyphlopidae, Epictinae, Epictini, Typhlopoidea, Serpentes, Squamata (snakes) 
Subspecies 
Common Names 
SynonymEpictia vindumi WALLACH 2016: 289
Glauconia albifrons — BARBOUR & COLE 1906: 150
Glauconia albifrons — FOWLER, 1913: 173
Glauconia albifrons — WERNER 1917: 203 (part)
Glauconia albifrons — HAAS 1930: 59, 73, figs. 90–92 (part)
Glauconia albifrons — HAAS 1931: 127 (part)
Glauconia albifrons — SHATTUCK 1933: 576
Glauconia albifrons — DUNN 1944: 80
Glauconia albifrons — SMITH & SMITH 1976: S-B-97, S-F-50.
Stenostoma albifrons — SCHMIDT 1936: 168
Leptotyphlops albifrons — GAIGE 1936: 298; 1938: 297
Leptotyphlops albifrons — SCHMIDT 1936: 168–169, 183
Leptotyphlops albifrons — SCHMIDT & ANDREWS 1936: 168ff
Leptotyphlops albifrons — BARRERA 1962: 9, 1963: 86
Leptotyphlops albifrons — SMITH & SMITH 1976: S-B-127ff.
Leptotyphlops goudoti phenops — SMITH & SMITH 1976: S-B-129–130 (part)Leptotyphlops goudoti phenops — CONANP 2012: 155.
Leptotyphlops goudotii phenops — PETERS et al. 1970
Leptotyphlops goudotii phenops — PETERS et al. 1986: 170 (part)
Leptotyphlops goudotii phenops — SMITH & SMITH 1976: S-B-129ff
Leptotyphlops goudotii phenops — HAHN 1980: 16 (part)
Leptotyphlops goudotii phenops — LEE 2000: 259.
Leptotyphlops phenops — SMITH 1939: 28
Leptotyphlops phenops — KLAUBER 1940: 151 (part)
Leptotyphlops phenops — MARTIN DEL CAMPO in PEARSE 1945: 225
Leptotyphlops phenops — LIST 1966: 90–91, 107, pl. 16, fig. 7
Leptotyphlops phenops — LANGEBARTEL 1968: 110ff, fig. 2K
Leptotyphlops phenops — SMITH & SMITH 1976: S-B-130ff (part)
Leptotyphlops phenops — LEE 1977: 146ff, fig. 43 (lower right)
Leptotyphlops phenops — LEE 1980: 11ff, pl. 15
Leptotyphlops phenops — FERRI 2000: 105 (part)
Leptotyphlops phenops phenops — SMITH & TAYLOR 1945: 24 (part)
Leptotyphlops phenops phenops — SMITH & SMITH 1976: S-B-130ff (part)
Leptotyphlops goudoti — HAHN 1979: 230.2–3 (part)
Leptotyphlops goudoti — FLORES-VILLELA 1993: 35 (part)
Leptotyphlops goudoti — WILSON & MCCRANIE 1998: 18, 31, 35 (part).
Leptotyphlops goudotti (sic) — VILLA et al. 1988: 85 (part).
Leptotyphlops goudotii — LAZCANO-BARRERA et al. 1992: 36
Leptotyphlops goudotii — HEDGES 1996: 111, 115 (part)
Leptotyphlops goudotii — LEE 1996: 280–281, fig. 151A, map (part)
Leptotyphlops goudotii — LEE 2000: 258–259, fig. 125A, col. fig. 306, map (part)
Leptotyphlops goudotii — CAMPBELL 1998: 186–187 (part)
Leptotyphlops goudotii — WALLACH 1998a: 490 (part)
Leptotyphlops goudotii — CUNDALL & IRISH 1998: 597
Leptotyphlops goudotii — LEE 2000: 258–259, fig. 125A, col. fig. 306, map 101
Leptotyphlops goudotii — SASA MARÍN 2000: 150 (part)
Leptotyphlops goudotii — KÖHLER 2001b: 12, fig. 11B (part)
Leptotyphlops goudotii — KÖHLER 2008: 183–184, map (part)
Leptotyphlops goudotii — KEARNEY 2002: 735
Leptotyphlops goudotii — GARCÍA-GIL 2006a: 220, 2006b: 209
Leptotyphlops goudotii — GONZÁLEZ-BACA, 2006: 13
Leptotyphlops goudotii — MEREDIZ-ALONSO et al., 2007: 89
Leptotyphlops goudotii — ELVIRA-QUESADA et al., 2007: 37, 89
Leptotyphlops goudotii — CONABIO 2009: 17 (part)
Leptotyphlops goudotii — ANONYMOUS 2010b: 38
Leptotyphlops goudotii — CHABLÉ-SANTOS 2010: 261, Anexo XV
Leptotyphlops goudotii — JOHNSON et al. 2010: 368 (part)
Leptotyphlops goudotii — WILSON et al. 2013: 44 (part)
Leptotyphlops goudotii — GUERRERO-GONZÁLEZ 2014: 2.
Epictia goudotii — CANSECO-MÁRQUEZ & GUTIÉRREZ-MAYÉN 2010: 252 (part). Epictia phenops — WALLACH et al. 2014: 278 (part). 
DistributionMexico (Yucatán, Quintana Roo), elevation 0-30 m.

Type locality: Chichén Itza, Yucatán, Mexico, 20°41'03"N, 88°34'06"W, elevation 35 m.  
Reproduction 
TypesHolotype: FMNH 153536 (field number EWA 79), a 153 mm (LOA) specimen collected by E. Wyllis Andrews in February of 1959.
Paratypes (45): MEXICO: YUCATÁN: Chichén Itza: collected by Leon J. Cole between 13 February and 9 April 1904, MCZ R-7113; collected by Oliver G. Ricketson, Jr., in 1925, MCZ R-22060–62; collected by Joseph C. Bequaert in June of 1929, MCZ R-28747; collected by E. Wyllis Andrews on 2 April 1934, FMNH 20616–17; collected by E. Wyllis Andrews on 5 April 1934, FMNH 20618; collected by E. Wyllis Andrews on 2 April 1939, FMNH 36337 (field no. EWA 610); collected by E. Wyllis Andrews on 1 October 1939, FMNH 36338 (field no. EWA 15), FMNH 36343 (field no. EWA 7); collected by E. Wyllis Andrews on 5 October 1939, FMNH 36344 (field no. EWA 53); collected by E. Wyllis Andrews on 17 October 1939, FMNH 36339 (field no. EWA 210); collected by E. Wyllis Andrews on 7 November 1939, FMNH 36340 (field no. EWA 494); collected by E. Wyllis Andrews on 18 November 1939, FMNH 36336 (field no. EWA 522); collected by E. Wyllis Andrews on 15 March 1940, FMNH 36332 (field no. EWA 1322); collected by E. Wyllis Andrews, date unknown, FMNH 36334 (field no. EWA 1295), FMNH 36335 (field no. EWA 1292). Dzibilchaltun: collected by E. Wyllis Andrews in December of 1958, FMNH 153532 (field no. EWA 33), FMNH 153533 (field no. EWA 33), FMNH 153535 (field no. EWA 46); collected by E. Wyllis Andrews in January of 1959, FMNH 153542 (field no. EWA 58), FMNH 153546 (field no. EWA 184); collected by E. Wyllis Andrews in February of 1959, FMNH 153543 (field no. EWA 74); collected by E. Wyllis Andrews in September of 1959, FMNH 153545 (field no. EWA 100); collected by E. Wyllis Andrews in April of 1960, FMNH 153501 (field no. EWA 139), FMNH 153537 (field no. EWA 135), FMNH 153544 (field no. EWA 137), FMNH 153596 (field no. EWA 138); collected by E. Wyllis Andrews in November of 1961, FMNH 153586 (field no. EWA 165); collected by E. Wyllis Andrews in January of 1962, FMNH 153538 (field no. EWA 192); collected by E. Wyllis Andrews in November of 1963, FMNH 153587 (field no. EWA 229), FMNH 153588 (field no. EWA 230), FMNH 153589 (field no. EWA 231). Kantunil: collected by E. Wyllis Andrews on 7 November 1939, FMNH 36341 (field no. EWA 496), FMNH 36342 (field no. EWA 495). Merida: Had Chenku: collected by E. Wyllis Andrews on 7 January 1942, FMNH 40724, collected by E. Wyllis Andrews on 20 January 1942, FMNH 40725; collected by Eduardo C. Welling on 10 March 1965, UCM 29978. Piste: collected by Eduardo C. Welling on 8 August 1967, CM 49549; collected by Eduardo C. Welling on 20 August 1967, CM 49550; collected by Eduardo C. Welling on 3 September 1967, CM 49551–52; collected by Eduardo C. Welling on 16 September 1967, CM 49553; collected by Eduardo C. Welling on 22 September 1967, CM 49554. 
CommentSynonymy: after Wallach 2016: 289. See this paper also for references.

Diagnosis: (1) scale row formula = 14-14-14; (2) midtail scale rows = 10; (3) total length = 72–163 (x– = 142.8) mm; (4) total midodrsals = 230–253 (x– = 243.6); (5) subcaudals = 15–23 (x– = 17.8); (6) relative body propor- tion = 40–70 (x– = 53.6); (7) relative tail length = 5.1%–9.0% (x– = 6.5%); (8) relative tail width = 2.8–5.7 (x– = 3.8); (9) relative rostral width = 0.24–0.44 (x– = 0.37); (10) relative eye size = 0.43–0.53 (x– = 0.48); (10) rostral parallel with a rounded apex, extending beyond the eye level; (11) supralabials 2, moderate anterior supralabial reaching mid-eye level; (12) frontal hexagonal, as broad as deep; (13) supraoculars large and pentagonal, twice as broad as deep, posterior border soblique to supranasal borders; (14) widest anteriormost vertebral scale 3rd; (15) parietals and occipitals subequal, oriented transversely; (16) infralabials 4; (17) cloacal shield semilunate in shape; (18) head brown, with a yellow spot covering entire rostral; (19) dorsum with 7–9 dark brown stripes of diamond-shaped spots, bordered by moderate zigzag yellow stripes (= 7–9 dark stripes); (20) venter pale brown, many scales out- lined in yellow; (21) midbody stripe formula (7–9 + 0) and middorsal pattern (3); (22) tail with a pale terminal spot covering dorsal scales 1–3 (x– = 1.7) and ventral scales 8–13 (x– = 9.8) (ventral/dorsal ratio 5.8); and (23) apical spine a small compressed spike.

Habitat: low scrubby deciduous thorn forest, ranging in height from three to seven meters and dominated by deciduous legumes (Lee, 1977).

Comparisons: see Wallach 2016: 294 for detailed comparisons. 
Etymology“This species is named in honor of Jens Vindum, Senior Collection Manager, Department of Herpetology, California Academy of Sciences (CAS), and commemorates his 35 years of service to the museum. During my early visits to the Academy from 1983 to 1985, to examine snake viscera, Jens was a most accommodating host during my stay, and since then always has lent material and shared data for my research.” 
References
  • Wallach, Van. 2016. Morphological review and taxonomic status of the Epictia phenops species group of Mesoamerica, with description of six new species and discussion of South American Epictia albifrons, E. goudotii, and E. tenella (Serpentes: Leptotyphlopidae: Epictinae Mesoamerican Herpetology 3 (2): 216-374 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
 
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