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Eremias scripta (STRAUCH, 1867)

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Higher TaxaLacertidae, Eremiadinae, Sauria, Lacertoidea, Squamata (lizards)
SubspeciesEremias scripta lasdini (TZAREVSKY 1918)
Eremias scripta pherganensis SZCZERBAK & VASHETKO 1973
Eremias scripta scripta (STRAUCH 1867) 
Common NamesE: Sand Racerunner 
SynonymPodarces (Scapteira) scripta STRAUCH 1867
Scapteira scripta — BOULENGER 1887: 112
Scapteira scripta — BOETTGER 1888
Scaptira [sic] scripta — ALCOCK & FINN 1897: 559
Eremias (Rhabderemias) scripta — LANTZ 1928: 90
Eremias scripta — SMITH 1935: 386
Scapteira bilkewitschi NIKOLSKY 1905
Scapteira grum-grzimailoi NIKOLSKY 1915 (part.)
Scapteira ladzini TZAREVSKY 1918
Rhabderemias scripta — LANTZ 1928
Eremias (Rhabderemias) scripta — BISCHOFF 1978
Rhabderemias scripta — DAS 1996: 49
Eremias scripta — LAMBERT 2002
Eremias scripta — SZCZERBAK 2003
Eremias (Rhabderemias) scripta — SINDACO & JEREMČENKO 2008: 232
Eremias scripta — NASRABADI et al. 2017

Eremias scripta lasdini (TZAREVSKY 1918)
Scapteira lasdini TZAREVSKY 1918: 84
Scapteira bilkewitschi NIKOLSKY 1905:486
Eremias scripta lasdini — BORKIN & DAREVSKY 1987
Eremias (Rhabderemias) scripta lasdini — VEDMEDERYA et al. 2009
Eremias lasdini — ORLOVA et al. 2023

Eremias scripta pherganensis SZCZERBAK & WASHETKO 1973
Eremias scripta pherganensis — BORKIN & DAREVSKY 1987
Eremias scripta pherganensis — BONDARENKO 2020
Eremias scripta pherganensis — ORLOVA et al. 2023 
DistributionS Kazakhstan, Turkmenistan, NE Iran, Afghanistan, NW Pakistan, SE Uzbekistan, SW Tajikistan, Fergana Valley in Uzbekistan, Kyrgyzstan

Type locality: Transcaspia; “Aralo-Caspian desert” fide ANDERSON 1999.

lasdini: SE Uzbekistan

pherganensis: Fergana Valley  
TypesLectotype: ZISP (ZIL) 3669 (designated by Szczerbak, 1974:213)
Holotype: ZISP 12107, Uzbekistan, Surxondaryo Region, Sands Katta-Kumy, vicinity Termez, 18.05.1915, Coll. V.J. Lazdin [lasdini]
Holotype: NMNHU (Zoological Museum named after N.N. Szczerbak National Museum of Natural History, National Academy of Sciences of Ukraine, Kiev) Re 6, Uzbekistan, Fergana Valley, Ak-Kum sands, 35.05.1969, Coll. E.V. Vashetko [pherganensis] 
DiagnosisDiagnosis: Subocular borders mouth; lateral scales of fourth toe in complete row for entire length of toe, forming distinct fringe, 2 complete rows of subdigital scales, i.e., total of 4 scales counted around fourth toe (except extra scale may be present atjoints) (Anderson 1999: 224, Fig. 108). 
CommentSynonymy partly after KHALIKOV & ANANJEVA (pers. comm.) 
  • Alcock, A. W. & FINN,F. 1897. An account of the Reptilia collected by Dr. F.P. Maynard, Captain A.H. McMahon, C.I.E., and the members of the Afghan-Baluch Boundary Comission of 1896. J. Asiat. Soc. Bengal 65: 550-566 [1896] - get paper here
  • Ananjeva, NB, Orlov NL; Khalikov RG; Darevsky IS; Ryabov IS & Barabanov AV 2006. The Reptiles of North Eurasia. Taxonomic Diversity, Distribution, Conservation Status [this comprises the territory of the former Soviet Union and Mongolia]. Pensoft Series Faunistica 47, 250 pp.
  • Anderson, Steven C 1999. The lizards of Iran. Contributions to Herpetology Volume 15, Society for the Study of Amphibians and Reptiles, Saint Louis, Missouri: i-vii, 1-442 [review in Copeia 2000 (4): 1144] - get paper here
  • Baig, Khalid Javed and Rafaqat Masroor 2006. A New Species of Eremias (Sauria: Lacertidae) from Cholistan Desert, Pakistan. Russ. J. Herpetol. 13 (3): 167-174
  • Bischoff, W. 1978. Beiträge zur Kenntnis der Echsen des Kaukasus. Salamandra 14 (4): 178-202 - get paper here
  • Bondarenko D. A. 2020. Community of Reptiles in the Sandy Habitats of the Ferghana Valley (Uzbekistan) and the Endemic Species Conservation Problem. Current Studies in Herpetology 20 (1-2): 3 - get paper here
  • Bondarenko D. A., Ergashev U. H. 2022. Reptiles of the Southwestern Tajikistan desert plains: Spatial distribution, population density and communities structure. [in Russian] Current Studies in Herpetology 22 (1-2): 17 - get paper here
  • Borkin L. Ya., Darevsky I.S. 1987. List of amphibians and reptiles of the USSR fauna // Amphibians and reptiles of protected areas. [in Russian] Collection of scientific papers. Moscow: Central Research Institute of Glavokhoty of the RSFSR. Pp. 128-141
  • Boulenger, G. A. 1887. Catalogue of the lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575 pp. - get paper here
  • Brushko, Z.K. 1995. Lizards of desert regions of Kazakhstan. [in Russian] Kazakhstan National Academy of Sciences, Institute of Zoology, Almaty, 232 pp.
  • Chirikova M.A. 2021. Historical context and results of a study on Racerunners (Sauria, Lacertidae, Eremias) in Kazakhstan [in Russian] Proc. Inst. Zool., Kazakhstan, pp. 127-136 - get paper here
  • CHIRIKOVA, M. A., Y. A. ZIMA, M. V. PESTOV, AND V. A. TERENTYEV 2020. Biodiversity of the Herpetofauna of the Muyunkum Desert, Kazakhstan. Herpetological Review 51: 438-446.
  • Davletbakov, A. T.; L. A. Kustareva, D. A. Milko, A. N. Ostastshenko, S. S. Sagymbaev, S. A. Toropov, N. V. Trotchenko 2015. Genetical Fund Cadastre of Kyrgyz Republic. Vol. IV. Phylum Chordata (chordates). Bishkek, 128 p. + XXXVI color tables - get paper here
  • Dujsebayeva, T. N. (ed.) 2010. Short review of last changes in the checklist of amphibians and reptiles of Kazakhstan. [in Russian]. In: Dujsebayeva, T. N. (ed.) Herpetological Researches in Kazakhstan and adjacent countries, p. 37-52
  • Eremchenko, V.K. & Panfilov, A.M. 1999. Taxonomic situation of multiocellated racerunner of the Eremias multiocellata complex of Kyrgyzstan and neighbour China (Sauria: Lacertidae: Eremias). [in Russian] Science and New Technologies - Nauka i Novye Tekhnologii, Ser. Biol., 4: 112-124
  • Guo, Xianguang; Xin Daib, Dali Chen, Theodore J. Papenfuss, Natalia B. Ananjeva, Daniel A. Melnikov, Yuezhao Wang 2011. Phylogeny and divergence times of some racerunner lizards (Lacertidae: Eremias) inferred from mitochondrial 16S rRNA gene segments. Molecular Phylogenetics and Evolution 61: 400–412 - get paper here
  • Khan, M.S. 2003. CHECKLIST AND KEY TO THE LIZARDS OF PAKISTAN. Pakistan J. Zool. Suppl. Ser. (1): 1-25 - get paper here
  • Lambert, M.R.K. 2002. Preliminary observations on herpetofaunal diversity in the Almaty region, Southern Kazakhstan (September 1998). Herpetological Bulletin (79): 7-13 - get paper here
  • Lantz,L.A. 1928. Les Eremias de l’Asia Occidentale. Bull. Mus. Géorgie vols. 4/5: 136 pp. - get paper here
  • Leviton, Alan E. 1959. Report on a collection of reptiles from Afghanistan. Proc. Cal. Acad. Sci. 29 (12): 445-463 - get paper here
  • Martin T, Guillemin M, Nivet-Mazerolles V, Landsmann C, Dubos J, Eudeline R, and Stroud J. 2017. The herpetofauna of central Uzbekistan. Amphibian & Reptile Conservation 11(1) [General Section]: 93–107 - get paper here
  • Mishagina J.V. 2005. On trophic connections of sand racerunner from Repetek reserve. Probl. osvoenya pustyn, 2005, No 3: 45-49 [in Russian]
  • Mishagina J.V. 2008. Ecological groups of preys in Eastern Kara Kum psammobiont lizards, Eremias (Lacertidae) and Phrynocephalus (Agamidae). The Problems of Herpetology (Proceedings of the 3rd Meeting of the Nikolsky Herpetological Society). Saint-Petersburg, 2008, pp. 298-307 [in Russian] - get paper here
  • Mishagina, J. V. M 2007. On food habits of sand racerunner, Eremias scripta (Strauch, 1867) (Squamata, Lacertidae) from barchan sands of Eastern Kara Kum (Repetek reserve). Russ. J. Herpetol. 14 (1): 50-56 - get paper here
  • NASRABADI, REZA; NASRULLAH RASTEGAR-POUYANI, ESKANDAR RASTEGAR-POUYANI, AHMAD GHARZI 2017. A revised key to the lizards of Iran (Reptilia: Squamata: Lacertilia). Zootaxa 4227 (3): 431–443 - get paper here
  • Nikolsky, A. M. 1905. Herpetologia rossica. Mem. Acad. Imp. Sci., Phys.-Math. Classe, St. Pétersbourg, XVII (1): 1-518
  • ORLOVA, VALENTINA F.; ESKANDAR RASEGAR-POUYANI, KHOSROW RAJABIZADEH, HOSSEIN NABIZADEH, NIKOLAY A. POYARKOV, DANIEL A. MELNIKOV, ROMAN A. NAZAROV 2023. Taxonomic diversity of racerunners with descriptions of two new Eremias species (Sauria: Lacertidae) from Central Iran. Zootaxa 5369 (3): 336–368 - get paper here
  • Shestopal A., Amanov A., Ovezov T. 2021. An overview of reptiles from the Kaplankyr Nature Reserve and adjacent territories. [in Russian] Proc. Inst. Zool., Kazakhstan, pp. 105-126 - get paper here
  • Sindaco, R. & Jeremcenko, V.K. 2008. The reptiles of the Western Palearctic. Edizioni Belvedere, Latina (Italy), 579 pp. - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Strauch,A. 1867. Bemerkungen über die Eidechsen-Gattung Scapteira (Fitz.). Bull. Ac. Sc. St. Pétersb. 12: 313-328 or Mél. Biol. 4: 403-426 (after ZR 1868: 116) - get paper here
  • Szczerbak, N.N. 2003. Guide to the Reptiles of the Eastern Palearctic. Krieger, Malabar, FL, 260 pp. - get paper here
  • Szczerbak, N.N. & Vashetko, E.V. 1973. New subspecies – Eremias scripta pherganensis ssp. n. (Reptilia, Sauria) from Uzbekistan [in Russian] Vestnik zoologii, Journal of the Schmalhausen Institute of Zoology, National Academy of Sciences of Ukraine, Kiev, 1973 (1): 84-86
  • Tian, L., & Guo, X. 2022. Complete Mitochondrial Genomes of Five Racerunners (Lacertidae: Eremias) and Comparison with Other Lacertids: Insights into the Structure and Evolution of the Control Region. Genes, 13(5): 726 - get paper here
  • Tzarevsky, S.F. [surname also translated as Carevskij, Czarevsky, Tsarewsky and Zarevskij fide Adler, 2012: 216] 1918. Reptiles et amphibies, collectionnés par V. J. Lazdin dans la Boukharie méridionale et orientale ainsi que dans la partie N.-E. de la Province Transcaspienne en été 1915. [in Russian] Annuaire Musée Zoologique de l’Académie Impériale des Sciences de St.-Pétersbourg, 22: 79-90 - get paper here
  • Vedmederya, Valeriy; Oleksandr Zinenko, and Andrei Barabanov 2009. An Annotated Type Catalogue of Amphibians and Reptiles in the Museum of Nature at V. N. Karazin Kharkiv National University (Kharkiv, Ukraine). Russ. J. Herpetol. 16 (3): 203-212 - get paper here
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