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Eutrachelophis steinbachi (BOULENGER, 1905)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Common Names 
SynonymRhadinaea Steinbachi BOULENGER 1905: 454
Aporophis melanocephalus GRIFFIN 1915: 171
Liophis steinbachi — AMARAL 1929: 174
Liophis steinbachi — VANZOLINI in PETERS & OREJAS-MIRANDA 1986: 16
Erythrolamprus steinbachi — GRAZZIOTIN et al. 2012
Rhadinaea steinbachi — WALLACH et al. 2014: 834
Eutrachelophis steinbachi — MYERS & MCDOWELL 2014 
DistributionSE Bolivia

Type locality: Provincia Sara, Dep. Santa Cruz de la Sierra, Bolivia  
TypesLectotype: BMNH 1946.1.21.62 (designated by MYERS & MCDOWELL 2014). Paralectotype: BMNH 1946.1.21.63 
DiagnosisDIAGNOSIS: Eutrachelophis steinbachi is like E. bassleri in having 15 dorsal scale rows and a usually conspicuous pair of ocellar markings6 on the nape. It is a larger snake (to 558 mm total length) than bassleri (, 400 mm) and is readily differentiated by details of color pattern. E. steinbachi has a pair of oblique pale markings touching the upper anterior and upper posterior edges of the eye (both lacking in bassleri), and the dark head color extends onto the neck as unbroken dorsal and lateral stripes. E. steinbachi lacks the broken, ocellarlike nuchal collar (present in bassleri). A lateral line of pale dashes, where present, lies on scale row 6 in steinbachi, on row 4 in bassleri. E. steinbachi differs absolutely in having a divided hemipenis with spinose tips [MYERS & MCDOWELL 2014: 16] 
CommentSynonymy that of PETERS & OREJAS-MIRANDA 1970. Incertae sedis fide DIXON 1980. 
Etymologynamed after Dr. Jose Steinbach (1856-1929), a botanical and zoological collector in Argentina and Bolivia for the Field Museum. 
  • Amaral,A. do 1930. Estudos sobre ophidios neotropicos XVIII. Lista remissiva dos ophidios da região neotropica. Mem. Inst. Butantan 4: 126-271 [1929] - get paper here
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, George A. 1905. Descriptions of new snakes in the collection of the British Museum. Ann. Mag. nat. Hist. (7) 15 (89): 453-456 - get paper here
  • Dixon, J. R. 1980. The neotropical colubrid snake genus Liophis. The generic concept. Milwaukee Public Museum Contributions in Biology and Geology 31: 1-40
  • Grazziotin, Felipe G.; Hussam Zaher, Robert W. Murphy, Gustavo Scrocchi, Marco A. Benavides, Ya-Ping Zhang and Sandro L. Bonatto 2012. Molecular phylogeny of the New World Dipsadidae (Serpentes: Colubroidea): a reappraisal. Cladistics DOI: 10.1111/j.1096-0031.2012.00393.x - get paper here
  • Griffin, L.F. 1916. A catalog of the Ophidia from South America at present (June 1916) contained in the Carnegie Museum with descriptions of some new species. Mem. Carnegie Mus. 7 (3) [1915]: 163-228 - get paper here
  • Myers, Charles W. and Samuel B. McDowell 2014. New Taxa and Cryptic Species of Neotropical Snakes (Xenodontinae), with Commentary on Hemipenes as Generic and Specific Characters. Bulletin of the American Museum of Natural History 385 (1) : 1-112. - get paper here
  • Valencia, J. H., K. Garzón-Tello & M. E. Barragán-Paladines 2016. Serpientes venenosas del Ecuador: sistemática, taxonomía, historia natural, conservación, envenenamiento y aspectos antropológicos. Quito, Ecuador, Fundación Herpetológica Gustavo Orcés, Universidad de Texas, Fondo Ambiental Nacional, 652 pp. [review in HR 49 (1): 152, 2018]
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
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